204 BOTANICAL GAZETTE [OCTOBER 
the ascogonium and ascogenous hyphae. The spindles of this 
division are similar to those of the first, and usually lie in a plane 
which is approximately parallel to the axis of the ascus, but, as 
HARPER (21) has shown, they may vary markedly from this 
position. At metaphase the five chromosomes divide and five pass 
to each pole. Telophase and the reorganization of the daughter 
nuclei appear to be entirely similar to the same processes as 
described at the end of the first division. 
The third division is essentially like the second, except that 
the spindles are usually approximately at right angles to the axis 
of the ascus although, as 
HarPER (21) has shown, 
one of them may be par- 
allel to the ascus wall. 
At telophase, when the 
masses of chromosomes 
have broken through 
the nuclear membrane, — 
some of the fibers which 
radiate from the centro- 
some out into the cyto- 
plasm appear to be con- 
nected to the plasma 
membrane around the 
Fics. 49-51.—Telophase of: third division; radi- Wh 
ating fibers attached to plasma membrane; fig. 50, — (fig. 49) : is 
nucleus reorganized after third division; fig. 51, this occurs, the plasma 
spore showing beginning of secondary thickening membrane is pulled in 
of wall; fibers still apparent; all X 2800 
° toward the groups of 
chromosomes as though the fibers which connect the groups of 
chromosomes with the plasma membrane, by contracting, were 
drawing the membrane and group of chromosomes together (fig. 
49). As the groups of chromosomes approach the periphery of 
the ascus, the radiating fibers come to be bent backward; this 
may be due to the movement of the centrosomes. The nuclei 
reorganize in a manner similar to that described for the daughter 
nuclei at the end of the first division, except that a more pro- 
nounced beak is formed on the nucleus where the radiating fibers 
