330 BOTANICAL GAZETTE [NOVEMBER 
al organs of both sexes, and because (2) self-fertilized hermaphro- 
dites produce dimorphic progenies, consisting of females and 
hermaphrodites. 
In my first paper on the inheritance of sex in Lychnis (SHULL 26), 
I represented the sex genes by the conventional signs for the sexes 
(9, 6, and ¥). As these signs were used in my tables with two 
different meanings—to represent sometimes the character of the 
genes and at other times the character of the soma—lI suspect that 
readers may have experienced some difficulty in comprehending 
the tables. I shall therefore adopt here the plan usually followed 
by students of genetics, of representing the genes by letters, letting 
Ff be respectively the presence and absence of a female determiner, 
Mm a male determiner, and Hk a hermaphrodite determiner. 
The conventional signs for the séxes will be used in this paper 
only in their more usual signification, referring to the nature of 
the soma, that is, the sporophyte. 
If Correns’ view of sex determination is correct, and the males 
are heterozygous, the females must be homozygous. CastTLe (5) 
suggests that in such a case the females will always be positive 
homozygotes, having a pair of sex genes (FF) corresponding with 
a single equivalent gene (Ff) in the male. I do not believe that 
this view can be substantiated, as there seems no good reason why 
females should not be negative homozygotes in some plants and 
animals, “neutral”? homozygotes in others, and positive homozy-- 
gotes in a third class. If the females are positive homozygotes, the 
somatic formula of the two sexes may be represented thus: FF=3, 
and Ff= 4; if the females are negative homozygotes, the correspond- 
ing symbols will be FFmm=, and FFMm=<é; and if the female 
is a “neutral” homozygote, the formulae of the two sexes will be 
FF=$%,and FM=4. Only the first two of these assumptions cor 
cerning the nature of the females were considered in my earlier 
paper, and either was found capable of explaining the results secured 
in the first generation, provided the presence of a partially jnde- 
pendent hermaphrodite factor (H) might also be assumed. 
Whether there was any genetic relationship between the her- 
maphrodites A and B which produced hermaphrodite offspring, and 
C and D which produced males, could not be determined in the first 
