356 BOTANICAL GAZETTE [NOVEMBER 
will be discussed later in connection with the nature of the 
hermaphrodites. 
Correns (6, p. 17), with undoubted justification, maintained 
that the germ cells of monoecious, hermaphrodite, and dioecious 
species possess the tendency to develop into individuals having 
the distribution of sex organs characteristic of the particular 
genotype to which they belong; but when he likens the association 
of organs of both sexes in the same individual to the mosaic of red 
and white colors in striped flowers, and of pigmented and white 
spots in the coats of spotted animals, his justification becomes less 
obvious. Both striped flowers and spotted pelages are known 
from many investigations to be due to the presence or absence of 
a definite Mendelian gene, a so-called “spotting factor” or “pat- 
tern factor.” ae 
One of the chief aims in the arrangement of my cultures for 
1910 was to test the possible existence of such a mosaic or “pat- 
tern factor,” H, as a proximate cause of hermaphroditism m 
Lychnis, and the most striking result secured is the decisive mannet 
in which such a possibility is denied. The hermaphrodite character 
is not only incapable of reaching expression in the female* (as 
might be expected, since the female is homozygous), but it is also 
as a rule not transmitted through the egg cell to the male 
offspring. The males in the progeny of any cross agree in their 
sexual type with the male parent of that cross, regardless of its 
antedecent history. All the assumptions and implications involved 
in the first section of table I, in which an independent gene H was 
postulated, may therefore be rejected. 
SI refer here only to the normal functional hermaphroditism with which this 
paper deals, and not the pseudo-hermaphroditism which results when females of 
Lychnis dioica are attacked by the smut, Ustilago violacea, as reported by STRAS- 
and (0) that male plants may be infected also, but such infection does not in this case 
result in the development of the female organs. 
