506 BOTANICAL GAZETTE [JUNE 
size and numerous and large archegonia are offset by the late devel- 
opment of walls and their persistent delicacy, by the apparent lack 
of a ventral canal cell, and by the rather specialized necks of the 
archegonia. Probably it presents more resemblances to the gameto- 
phytes of the Taxaceae and to those of the Taxodineae than to 
other conifers. 
Summary 
1. The ovule possesses a very free nucellus with a glandular tip, 
a single integument adherent to the scale for almost its entire 
length, a ligule, a-_large micropyle, and spongy tissue surrounding 
the gametophyte. 
2. There is probably a single functional megaspore, which 
develops into an embryo sac with about 2000 free nuclei before 
cell-formation. 
3. Cell-formation follows on a peculiar centripetal growth of the 
cytoplasm and precedes wall-formation. 
4. The first walls are formed on the surface of the free cells. 
5. Secondary walls are formed on the spindles of the mitoses 
occurring in the primary cells of the peripheral regions of the 
gametophyte. 
6. The outer cells are uninucleate, the inner ones are multi- 
nucleate. 
7. The archegonia have single-tiered necks, usually, consisting 
of about 12 wedge-shaped cells. 
8. The necks are on the surface of the prothallus but are often 
overgrown. 
g. The archegonia may be single or occur in complexes and have 
a single-layered jacket. 
ro. A ventral canal nucleus may be absent, 
STANFORD UNIVERSITY 
CALIFORNIA 
LITERATURE CITED 
1. BURLINGAME, L. LANCELOT, The morphology of Araucaria brasiliensis. 1. 
The staminate cone and male gametophyte. Bor. Gaz. 55:97-114. pls. 
, 5 1914. 
a. Cee. J. M., and Lanp, W. J. G., The gametophytes and embryo of Tor- 
reya taxifolia. Bort. GAZ. 39:161-178. pls. 1-3. 1905. 
