84 BOTANICAL GAZETTE | AUGUST 
rise to the term “stalk cell.” It will be remembered that the division 
takes place in the former before and in the latter after germination 
has begun. Fig. 18 is a diagram showing this stage in Dioon, taken 
from CHAMBERLAIN (3). If in the case of Phyllocladus the genera- 
tive cell should enlarge upward toward the tube before the division, 
we can easily see that the oblique wall, failing to touch the prothallial 
cell, would give us the Dioon situation. On the other hand, if the 
wall in Dioon were a little lower down or slightly more oblique, we 
should have a section very like that of Phyllocladus. From this 
point of view the distinction between the anticlinal and periclinal 
division appears to be related to the time at which the division takes 
place, and to be of little significance in itself. 
Throughout the development of the gametophyte, though distinct 
cells are formed, each bounded by a Hauéschicht, there is no evidence 
of cellulose walls. This I believe to be true also in the Dacrydium 
and Podocarpus which I have examined. Nor&én (11) fails to find . 
cellulose walls in the case of Saxegothaea except in the first prothallial 
cell, and THompson (17) finds them in Microcachrys only in the 
prothallial cells. 
Miss KitpaHt mentions the occasional persistence of the first 
prothallial cell. Figs. 11 and 12 show not only this but also a still 
more rare case in which the second has divided. This, it will be 
remembered, is the usual condition in Dacrydium, Microcachrys, and 
Saxegothaea. In Dacrydium, moreover, the first prothallial cell very 
often degenerates early, which makes the resemblance to Phyllocladus — 
still stronger. 
As in all the Podocarpineae so far studied, the prothallial and — 
stalk nuclei become free in the general cytoplasm. This may occ , 
in Phyllocladus before the grains are shed, but sometimes not until — 
after they reach the micropyle. The mature grain contains the body 
cell, and the free prothallial, stalk, and tube nuclei. 
Young tubes were found in the nucellus November 20. Fig. 19 3 | 
shows the tube nucleus in advance, followed by the others, and thé: 
distinctly organized body cell still in the grain. 
My work does not confirm Miss KinpaAnt’s in regard to the male : 
cells, as in every case where a complete series of sections was 
secured, a decided difference in size was evident. The functional 
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