98 BOT. ANICAL GAZETTE [AUGUST 
The greatest gap in our present knowledge of conifers is shown — 
by the Araucarineae. Even in the male gametophyte, which supplies 
perhaps the strongest argument in the above discussion, a good deal 
of work is needed. The female gametophyte is little known and 4 
the embryo practically not at all. Beside this, we should know the — 
early development of the ovulate strobilus for a proper understanding 
of the morphology of its parts. In the Podocarpineae we lack 4 
adequate knowledge of the female gametophyte, embryo, and develop- _ 
ment of ovulate structures. Until further data on these points are 
available, we should be hardly justified in coming to a definite deci- 
sion in regard to relationships, and at present it seems best to hold 
Taxineae, Podocarpineae, and Araucarineae apart as separate tribes, 
leaving open the question of larger groupings among conifers. 
Grateful acknowledgments are due to Professors J. M. COULTER 
and C. J. CHAMBERLAIN, under whose direction this work was done. 
THE UNIVERSITY OF CHICAGO 
LITERATURE CITED 
1. Brooks, F. T., and Stmes, W., The structure of Podocarpus spinulosa 
(Smith) R. Br. Annals of Botiaiy 24: 305-318. pl. 21. 1910 
BurincamE, L. L., The staminate cone and male camstophyt of Podo- 
a 
carpus. Bor. Gin 46:161-178. pls. 8, 9. 1908. 
3. CHAMBERLAIN, C. J., Spermatogenesis in Dioon edule. Bot. GAZETTE 
— -473215-236. pl. 15. 1909. 
4. Coxer, W. C., Fertilization and embryogeny in Cephalotaxus Fortunet. 
Bot. GAZETTE 43:1-10. pl. 1. 1907. : 
5. —--—, Notes on the gia sew and embryo of Podocarpus. Bot. 
Gases 33:89-107. pls. 5-7. . 
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taxijolia. Bor. GAZETTE 39:161-178. pls. A, 1-3. 1905. : 
7. JEFFREY, E. C., and Curyster, M. A., The sicibeniaetophyte of the Podo- 
carpineae. Amer. Nat. 41:355-364. 1907. 
8. Kivpant, N. J., The morphology of Phyllocladus alpina. Bot. GAZETTE 
46: 339-347. pls. 20-22. 1908. 
‘ , Affinities of Phyllocladus. Bort. GazettE 46:464-465. 1908. 
Lopnrore, G., Ueber die Vielkemigkeit der Pollenkérner und Pollen 
schlauche von Aveacavin Bidwillii. Ber. Deutsch. Bot. Gesell. 23: 335-340: 
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