444 BOTANICAL GAZETTE [DECEMBER 
ole 
They appear, however, to be connected with similar hyphae which 
become very distinct only a little farther up (about 10) in the 
ascocarp. ‘These latter hyphae, of which there are three or four, 
are the ascogenous hyphae. They are very much larger than the 
surrounding vegetative hyphae, and in the character of their con- 
tents are quite distinct from them. The ascogenous hyphae 
become more and more distinct as they get nearer the tip of the 
ascocarp, but they show little tendency to branch until they have 
reached about one-half the distance from the base to the tip, 
when they divide to form a large number of smaller, much branch- 
ing hyphae. These latter hyphae have a tendency to spread out 
as well as to grow upward, but in young specimens they are of 
course near the larger hyphae. Assuming that the large cell at 
the base of the ascocarp is an ascogonium, as seems to be the case, 
and that it is connected with the ascogenous hyphae, this stage 
may be represented diagrammatically as in fig. 7, in which the 
black lines represent the ascogenous hyphae. In slightly older 
specimens the ascogenous hyphae have spread over the entire 
tip. At about this time the tip enlarges to form the head of the 
ascocarp. Some of the vegetative hyphae soon grow out to form 
the paraphyses, after which the ascogenous hyphae give rise to 
asci. This stage is represented diagrammatically in fig. 8, in which 
the black lines still represent the ascogenous hyphae, the older 
parts of which have disappeared. 
While the hymenium is being differentiated, some of the vegeta- 
tive hyphae give rise to large storage cells (fig. 9). This process 
is similar in the two species studied and the same description will 
apply to both. These large storage cells are formed in rows and 
give rise to paraphyses (fig. 10). The storage cells are at first 
multinucleate, but the nuclei usually fuse as growth proceeds 
(figs. 11-14). This process continues until often the cells con-. 
tain a single, very large nucleus, many times the size of the largest 
nucleus in the ascus (figs. 15-17). Frequently the nuclei are very 
irregular (fig. 18). In other cases, a cell may contain a very large 
ntcleus with several much smaller ones lying near it (fig. 19)- 
These last two cases suggest amitoses, but may very well be stages 
in fusion. The storage cells may have a very irregular shape 
