1g2t] SCHERTZ—MOTTLING 113 
about 30 per cent, and those in F, had almost disappeared. As 
the main part of the phosphorus in F, probably existed as phospho- 
protein, there was apparently a hydrolysis of this going on to 
form hydrolytic products of nucleic acids. Since little or no 
phosphorus was found in F,, apparently the phosphatides had 
nearly all disappeared. Germination (40) in darkness appears to 
be correlated with a pronounced decomposition of phosphorus 
containing proteins. Apparently during the mottling of Coleus 
there is a similar phenomenon. PaAttaprn (40) showed that as 
TABLE XXVII 
PHOSPHATES 
Samplh Sample Sampl hosph 
Lest wot to ao no. fit | Average | Dertoo gm. 
dry weight 
Me Pe a 0.0330 0.0184 0.0213 ©,0242 0.526 
SON Fk ieee 0.0013 9.0008 0.0007 ©. 0009 °0,.068 
Wee fe ©.0027 0.0026 0.0030 0.0028 0.425 
Total dry weight...... ©.0370 0.0218 0.0250 0276. bio ews sss 
meted A Be ci 0.0068 0.0056 0.0064 0.160 
Mott Se of aed is 0.0018 0.0014 0.0018 0.0017 0.107 
Madre 4B a en None None None None None 
Total dry weight...... °©.0087 0.0082 0.0074 Sos a aS 
Mottled Be Fy. cians. ec: .0067 0.0095 0.0080 0.0081 0.178 
pte BB ee 2.0018 ©.0012 0.0012 0.0014 004 
paottied BOF. None 0.0010 0.0010 ° 7 0.212 
Total dry weight...... 0.0085 0.0017 0.0102 | ve ae ae er ies 
proteins decomposed the inorganic phosphates increased. From 
table XXVII it is apparent that the phosphoproteins were being 
rapidly decomposed as the leaves mottled. If the phospholipins 
are estimated according to CzAPEK (10), by multiplying the amount 
of magnesium pyrophosphate in the ether extract by 7.27, then 
0.073 per cent of the wet green leaves and 1.009 per cent of the dry 
weight is lecithin. In the mottled leaves the phospholipins have 
almost entirely disappeared. The ratio of phosphorus (table 
XXVII) to nitrogen (table XXVI) is about 1 to 7 in the green 
leaves, while in the mottled leaves the ratio is about 1 to 11. 
