39° BOTANICAL GAZETTE [MAY 
and surrounded by a band of meristem which continues the growth 
of the thallus and produces archegonia and antheridia. 
3. The gametophytes are typically monoecious. All bene 
archegonia and many antheridia also. Antheridia and archegonia 
occur without order on the thalli and often within a few cells of 
each other and of sporophytes. 
4. Antheridia and archegonia may continue to develop until 
one or more sporophytes have attained considerable size. 
5. Antheridia all develop as is characteristic of eusporangiate 
ferns, whether in a massive tissue or on a slender branch. 
6. Archegonia develop as is characteristic of the group. 
7. Sporophyte development is in agreement with common 
descriptions. The leaves of the first shoot are three or four in 
number, of the second shoot four or five. 
8. Gametophytes grow to maturity under simple methods of 
culture. 
The writer is indebted to Professor CHARLES J. CHAMBERLAIN 
for his interest and helpful suggestions during the progress of these 
studies, and for spores for culture growth; also to Mr. C. T. WALKER 
for spores sent from western Oregon, to Professor T. J. FitzPATRICK 
for the identification of sporophytes and for the careful reading of 
manuscript and proof, and to Mr. GrorcE SwALE, without whose 
efficient care of the greenhouse and cultures the experiments could 
not have been carried out. 
UNIVERSITY OF NEBRASKA 
LINcoLn, NEB 
LITERATURE CITED 
1. BUCHTIEN, OTTO, rat ein nr a des Prothallium von Equisetum. 
Bibliotheca Botanica 8. 1887. 
. CampBELt, D. H., The annie and development of mosses and ferns 
(Archegontataey, Naw York. 1895. 
3- GOEBEL, K., Organography of plants. Transl. byI.B.Batrour. Oxford. 
1900. 
N 
4. Fitzpatrick, T. J., The fern flora of Nebraska. Amer. Fern Jour. 10: 6-15, 
34-44. 1920 
5. Horaerster, W., The higher Cryptogamia. Ray. Soc. transl. by F. 
CURREY. 1862. 
