1922] BLOMQUIST—ANGIOPTERIS 183 
presence or absence of an endodermis, the origin of lateral roots, 
and whether the central strand is of cauline or foliar origin. The 
relationship between roots and leaves and the vascular arrange- 
ments in the stipules are also points still unsettled. 
Material and methods 
The material used in this investigation was collected on the 
island of Tutuila, Samoa, by Dr. W. J. G. LAnp, of the Hull Botani- 
cal Laboratory, during October and November 1912. It consisted of 
an abundance of young sporophytes of Angiopieris evecta, ranging 
from the first to the tenth leaf stages and a few somewhat more 
advanced. 
This material was imbedded in paraffin, and complete serial 
sections ranging from 10 to 20 win thickness were made with a 
rotary microtome. These sections were then studied from below 
upward, and diagrammatic reconstructions were drawn of the 
different stages. From these a final reconstruction was made in 
which the different stages were superposed one upon another to 
show as clearly as possible the successive stages which the plant 
passes through in the progress of its development. The most 
serious difficulty encountered was the fact that as the sporeling 
develops beyond the third and fourth stages, the lower part of the 
stem becomes more or less distorted, and in still older stages even 
decays. This difficulty was overcome by studying a close series of 
stages, which the material provided, selecting characteristic 
sections from the younger stages, and carefully matching these 
with sections representing the same morphological location in the 
older stages. The possibility of misinterpretation due to local 
variations, which are quite common in the anatomy of Angiopteris, 
was guarded against by studying several specimens of the same or 
approximately the same stage. 
Investigation 
EXTERNAL FEATURES 
While the external features of Angiopieris have been described 
by several investigators, these will be reviewed in part to assist in 
relating the internal structures with the external features. The 
