190 BOTANICAL GAZETTE [MARCH 
FIRST LEAF STAGE 
The first few stages have been described by FARMER (8) and 
CAMPBELL (2), and the present investigation adds little to their 
descriptions. The primary root is stated by them to be diarch. 
The writer found this to be true in some cases, but in as many cases 
the root was found to be triarch. Similar variations have been 
reported by CAMPBELL in Danaea, Botrychium, and Helminthostachys 
(3). The endodermis is well organized and is separated from the 
protoxylem and protophloem by a uniseriate pericycle. In some 
places the protoxylem abuts directly on the endodermis, as has 
been reported previously (8). In the diarch stele there are two 
protophloem points and in the triarch three (figs. 16, 17). 
The transition region in Angiopteris, as reported for other 
Marattiaceae, is a solid strand of vascular tissue with more or less 
irregular outl‘ne (fig. 19). In passing from this region upward, 
some cells in the xylem remain unlignified, usually near the center 
of the stele, so that the latter may become more or less tubular 
in cross-section. This condition has been called a solenostele 
(fig. 25). In some cases, however, these parenchyma cells are 
more scattered, and the condition resembles what has been reported 
for Danaea by BREBNER (1). In this genus the parenchyma cells 
appear without any relation to the center of the protostele. In 
either case the appearance of these unlignified cells marks the 
union of the next root stele and the corresponding leaf trace above. 
The writer agrees with BrEBNER that ‘“‘the parenchyma. . 
called pith is probably simply due to the delat ana and tie 
preparation of the departure of the leaf trace.” No phloem is 
developed in the center of the tubular stele at this stage. 
A longitudinal section of the sporeling at this stage shows a 
well defined apical cell at the stem tip (fig. 18). This apical cell 
has been reported by CAMPBELL in Macroglossum (5), by Miss 
CuHarLes in Marattia (7), and by West in Danaea (13). In a 
transverse section it was impossible to locate definitely this apical 
cell, but from the shape of the cells in the stem tip region it seems 
that this cell is four-sided. No procambial tissue is to be observed 
below the stem tip, a condition which has been reported by BREBNER 
for Danaea and by CAMPBELL for Angiopteris. The first differentia- 
