198 BOTANICAL GAZETTE . [MARCH 
7. The leaf traces at first are single, but later bifurcate in the 
stipular region, but anastomose beyond. The point of forking 
moves closer and closer to the central strand. Further forking and 
anastomosing of the leaf trace take place in the more advanced stages. 
8. One strand from each side of the leaf trace goes to the lobes 
of the stipule. Branches from this strand supply the stipular 
lobes with vascular tissue. 
g. All the stipular strands end blindly except the lowest or 
main strand, which is terminated by a procambial strand originating 
from a group of meristematic cells on the inner side of the edge of 
the stipule. This group of cells probably helps to build up the 
fleshy stipule. 
10. The absence of cauline procambium and the definite relation 
between roots and leaves suggest that the vascular tissue of the 
central region is a sympodium of leaf traces, and most if not all 
of the central strand is of foliar origin. 
The writer wishes to express his gratitude to Dr. W. J. G. LAND 
for the valuable collection of material furnished for this investiga- 
tion and for his advice, criticism, and encouragement during the 
progress of the work. 
TRINITY COLLEGE 
DurHaMm, N.C. 
LITERATURE CITED 
1. BREBNER, G., On the anatomy of Danaea and other Marattiaceae. Ann. 
Botany 16:517-552. pls. 22, 23. 1902. 
. CAMPBELL, D. H., The embryo and young sporophyte of Angiopteris and 
Kaulfussia. Ann. Jard. Bot. Buitenzorg 3:69-82. 1910 
, The eusporagiate ferns. Carnegie Publ. 140. rort. 
, The prothallium and embryo of Danaea. Preliminary note. 
Ann. Botany 23:691. 1 
, The structures and affinities of Macroglossum alidae Copeland. 
Ann. Botany 28:651-669. pls. 46-48. 1914. 
, The eusporangiate ferns and the stelar theory. Amer. Jour. Bot. 
8: 303-315. 1921 
. CHARLES, GRACE, The anatomy of the sporelings of Marattia alata. Bor. 
GAZ. 51:81-101. 1911. 
. Farmek, J. B., and Hitt, T. G., On the arrangement and structure of the 
vascular strands in Angiopteris evecta and some other Marattiaceae. 
Ann. Botany 16:371-402. pls. 16-18. 1902. 
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