1922] GARDNER & KENDRICK—TOMATO MOSAIC 473 
from the mosaic plants, developed the disease. On July 28, 1921, 
sixteen potted tomato plants in the greenhouse were similarly 
inoculated with the juice from mosaic P. heterophylla plants, and 
by August 18 nine had developed mosaic. Four uninoculated 
plants held as controls remained healthy. In a tomato field, on 
August 21, 1919, thirteen plants of P. subglabrata were inoculated 
with the juice from crushed leaves of mosaic tomato plants. 
Twelve days later eight had developed mosaic. Nearby uninocu- 
lated plants observed as controls did not develop the disease. 
Late in August 1919, seventeen potted tomato plants in the green- 
house were inoculated with the juice from mosaic P. subglabrata 
' plants, and fourteen developed the disease; the seventeen uninocu- 
lated control plants remaining free from mosaic. On May 25, 
1921, four potted tomato plants in the greenhouse were inoculated 
by wounding the stem with a needle and rubbing the wounded 
region with cotton soaked in the juice of crushed leaves of mosaic 
P. subglabrata collected at Frankfort, and fifteen days later all had 
developed mosaic. The two control plants, similarly treated except 
that distilled water was substituted for the mosaic virus, remained 
healthy. On July 15, 1921, forty-seven tomato seedlings grown 
under a cloth cage in the field were inoculated with the virus from 
mosaic P. virginiana. Ten days later twenty had developed 
mosaic. None of the numerous uninoculated seedlings in the 
cage developed the disease. The identity of this Physalis species | 
was verified by Paut C. STANDLEY of the United States National 
Museum. The results of these cross inoculations show that the 
mosaic disease found on these weeds in the field is transmissible to 
tomatoes. 
OBSERVATIONS ON PHYSALIS MOSAIC IN 1919 AND 1920 
The attention of the writers was directed to the importance 
of Physalis as a carrier of tomato mosaic in the summer of 19109. 
Large numbers of P. subglabrata occurred in an experimental field 
of tomatoes near Frankfort, Indiana. Mosaic became epiphytotic 
on the tomatoes during the latter part of the season, and also 
appeared on many of the Physalis plants, especially in a low-lying 
section of the field where the weeds were most abundant. In this 
