1910] SHULL—INHERITANCE OF SEX IN LYCHNIS 113 
For convenience, I will designate the four plants successfully 
used in breeding by: the letters A, B, C, and D. When A was self 
fertilized it produced, as a result of two different operations, 33 
_ females and 25 hermaphrodites. When its pollen was used to fertilize 
four different females, the resultant progenies consisted of 236 females, 
161 hermaphrodites, and 2 males. When castrated and fertilized 
with pollen from a normal male, A gave rise to a progeny of 21 females, 
2 hermaphrodites, and 11 males. B was also self fertilized and gave 
a progeny of 110 females and g5 hermaphrodites. When pollen of 
B was used to fertilize three different females, it produced 162 females 
and 144 hermaphrodites. It is thus seen that the two plants, A and 
B, showed identical behavior and together produced self fertilized 
offspring consisting of 143 females and 120 hermaphrodites, and 
when crossed with females gave a total of 398 females, 305 hermaph- 
rodites, and 2 males. In any explanation of these results the 
occurrence of these two males will probably have to be left aside as 
wholly exceptional. Only further breeding will show whether they 
were true males, or hermaphrodites with pistils suppressed, perhaps, 
by some cause external to the germ cells. Plants C and D gave quite 
a different result. Attempts to self fertilize them and to cross them 
with normal males all proved futile, though more persistent efforts 
perhaps might have succeeded. Both were used as pollen parents 
in crosses with normal females. In such a cross C gave a progeny 
of 39 females and 5 5 normal males, and D gave 26 females and 18 
normal males. The details of the several crosses are given in table 
I, p. 114. : 
Considering first plants A and B, and leaving out of consideration 
for the present the two males occurring in crosses between females and 
hermaphrodites, and the two hermaphrodites which appeared in the 
cross between a hermaphrodite and a normal male, it is apparent 
that the hermaphrodite character belongs only to the males, for in the 
families in which these hermaphrodites were the pollen parents, the off- 
spring always showed the same ratios of females and hermaphrodites 
that would have been expected of females and males, had a normal 
male been used as the pollen parent. 
It is clear that the hermaphrodite individuals, C and D, belong 
to an entirely different category from A and B, for in the families 
