202 BOTANICAL GAZETTE [MARCH 
Moreover, they are not traceable to any distinct source, as were those 
in Mitrula, but arise merely from the ordinary hyphal threads of the 
fruiting body. This is an agreement with BREFELD’s observations 
on several species of Morchella, Helvella, and Geoglossum, as well 
as Leotia lubrica and Peziza Sclerotiorum (Heft IV, 130). According 
to him the ascogenous hyphae may 
arise from the common filaments 
of the fruiting body and even from 
those which had formerly pro- 
duced paraphyses. 
In Helvella elastica there is 
frequently a thickening of the 
sterile filaments before they form 
the profusely branched mass of 
paraphyses, but such enlargements 
have no connection with those 
occurring in theascogenous hyphae. 
The cells of the former are short, 
bulbous, and usually binucleate, 
while in the fertile threads they 
] ‘ are long, uniform, and multinu- 
cleate (see text jfig.). Moreover, no 
connection can be traced between 
the two, nor can any paraphyses be 
S seen coming from the ascogenous 
fertile filament; c, ee fertile flament From the fertile subhymenial 
ine sitymenll lve; 6928 tayer Jateral branches are given 
the fusion of terminal and antepenulti- Off in rather scanty numbers, and 
mate cells, and the formation of the these grow toward the layer of 
ascus (the unshaded parts are empty). paraphyses, where they branch 
repeatedly. The end cells of these lateral branches contain an 
abundance of deeply staining protoplasm and 2-5 nuclei, the usual 
number, however, being 2 (jigs. 11-19). The whole energy of 
growth seems to be concentrated in this end cell, for usually 
those behind it are completely empty, and long before branching 
has been completed even the walls disappear. DitTRicH notes 
