466 BOTANICAL GAZETTE : [JUNE 
discovered by ScHIONNING;'® and Endomyces magnusii, discovered by 
Eremascus fertilis is described as having a branched, septate mycelium, the 
cells of which are two to four-nucleate. The cells which are about to produce the 
gametes become uninucleate by putting in septa. Isogamous conjugation results 
from the fusion of the contents of lateral diverticula from adjoining cells. At the 
point of fusion an eight-spored ascus is formed, in which spore formation is like 
that in the yeasts. Later parthenogenetic asci are formed, which after the usual 
nuclear divisions contain four or more spores, some of which may abort. € 
nuclei of the vegetative cells are so small that their divisions could not be observed. 
Endomyces fibuliger differs from Eremascus fertilis in having uninucleate cells, 
yeastlike conidia which bud off from the cells of the mycelium, and no fusion of the 
contents of the diverticula. The asci are always parthenogenetically formed in 
~ faster growing protuberance. The author thinks that we have here the 
ns of an ancestral conjugation absolutely identical with that in Eremascus 
jerilis. -Saccharomycopsis capsularis differs from Endomyces fibuliger in that the 
cells of the mycelium may become asci or these may bud off from the mycelial 
cells. Endomyces magnusii differs from the preceding forms in having no yeast- 
like conidia, but the cells of the mycelium readily separate to form oidia. Asci 
result from the fusion of the contents of uninucleate oogones and antheridia, 
which are formed at the ends of closely or diiatitty related hyphae. About one- 
fourth of the asci arise parthenogenetically. The author holds that this form is 
related to the Schizosaccharomycetes through its oidia. 
According to VAN TIEGHEM’s’? classification, the family Eremascaceae 
includes Eremascus, Endomyces, Saccharomyces, Podocapsa, Olinea, Protomyces, 
Ascoidea, and Dipodascus. GUILLIERMOND would place the last of these genera 
in a separate group because in these the asci arise as plurinucleate structures and 
form many (a variable number) spores. He would group the yeasts, Eremascus, 
and Endomyces together. 
DANGEARD*S thought that the gametes of Eremascus were multinucleate and 
has placed this genus with Dipodascus, but the author holds that SroPPEL’s 
work and his own show that Eremascus and Endomyces should be in the same 
group. The author criticizes ENGLER and PRANTL’s description of Eremascus, 
and says that it is no longer valid because isogamous conjugation with twisting 
of gametes is not present in all species. He would retain the genus Eremascus, 
including E. fertilis and E. albus, and characterize it by the absence of conidia 
10 SCHIONNING, H., Nouveau genre de la famille des Saccharomycetes. CR de 
traveaux du lab. Carlsberg 6:93-113. 1903. 
11 Lupwic, F., Ueber Alkoholgirung und Schleimfluss lebender Baume. Ber- 
Deutsch, Bot. Gesell. 4: Gen. Versammlungs-Heft. XVII-XXVII 
12 VAN TIEGHEM, PuH., Eléments de botanique. 1908. 
13 DANGEARD, P. A., L’origine du périthéce chez les ascomycetes. Le Botaniste 
9 and 10:1906. 
