

1 9 20] WA LKER—C YA TH US 9 



appears as a darker area (fig. 25). Later, just to the outside of 

 the inner wall of the peridiole, there appears, between the peridiole 

 and the peridium, a region of actively growing filaments which 

 take the stain readily (fig. 13; higher magnification of left hand 

 peridiole, fig. 28). The filaments in this region elongate rapidly, 

 and soon form a bundle of parallel filaments, as shown in fig. 30, 

 which is a higher magnification of the lower left hand peridiole 

 seen in figs. 15 and 18. Surrounding this bundle are the somewhat 

 gelatinized filaments of the ground tissue, which will form the outer 

 covering of the mature funiculus. The filaments which make up 

 the central bundle of the funiculus continue very active growth, 

 and, being confined by the gelatinizing filaments of the ground 

 tissue, become coiled irregularly in this part. The filaments of 

 the central cord always remain more or less parallel, even in the 



mature funiculus. 



In the mature funiculus (fig. 23) the central cord is attached 

 to the peridiole by the parallel filaments which marked its origin. 

 It immediately twists to form the central coil of the funiculus, 

 and from this passes abruptly at the base into a region of more 

 delicate filaments which merge with the now gelatinizing tissue on 

 the inner wall of the peridium. This central strand is surrounded 

 by the gelatinizing filaments of the ground tissue which constitute 

 its sheath. 



The attachment of the funiculus to the peridium is always very 



weak in this species, so weak, in fact, that as one examines older 



fruits casually it is not evident, because the peridioles do not seem 



to be attached. An examination of the peridiole itself, however, 



hows that the well developed funiculus is always present. 



Development of pekidium and epiphragm. — The mature 



peridium (fig. 33) is made up of three definite regions, a loosely 

 interwoven outer layer composed of largely longitudinal filaments 

 giving rise to hairs on the surface (a), a compact pseudoparenchy- 

 matic layer just w r ithin this (6), and a layer of loosely interwoven, 

 more or less gelatinized filaments which extend from the pseudo- 

 parenchymatic layer to the glebal region (c). 



In the development of the fruit body one can easily trace the 

 origin of these layers. The differentiation takes place first at 



