1920] WALKER— CYATH US 19 



funiculus is entirely lacking. Both are covered at first with 

 toothed hairs, and the internal structures are similar. The zone 

 of gelatinization, which is the first differentiation to be observed, 



similar 



liocarp 



basal, as shown by the location of the toothed hairs on the upper 

 portions of mature basidiocarps. The tissues covering the top 

 of the fruit body just before opening are seemingly homologous, 

 but more definitely limited in Crucibulum. The structure of the 

 peridium and the origin and development of the peridioles are 

 similar, but in Nidularia they appear first at the base, while in 

 Crucibulum they appear simultaneously. The structure of the 

 walls of the peridioles, as described by Fries, is very different 

 seemingly. The close relationship of these forms, however, is 

 very evident. 



more 



of the peridium and funiculus, but the general type of development 

 is very similar. Of the forms studied, C. fascicularis shows closer 

 resemblances to Crucibulum than does C. striatus. The part of 

 the gleba to become differentiated first and mature first seems to 

 be very variable in the genera and species studied. While in my 

 materials it has seemed to remain quite constant for each species, 

 there is the possibility that it may vary even in the same species. 



Summary 



1. All three species are easily grown on artificial media. Mature 

 fruit bodies were obtained only in cultures of Cyathus fascicularis. 



2. The mycelia of all are very similar except for color. Clamp 

 connections are abundantly present, and conspicuous mycelial 

 strands are formed. The cells of Cyathus fascicularis are binu- 

 cleate or composed of segments with paired nuclei. 



3. The basidiocarps of Cyathus fascicularis and C. striatus arise 

 from mycelial strands in all cases observed; while those of Cru- 

 cibulum vulgar e may arise from mycelial strand^, dense mats of 

 hvphae, or from the interior of old peridia. 



4. The primordium of the basidiocarp seems to have its origin 

 slightly below the tip of the strand, and consists of closely inter- 

 woven filaments smaller than those of the strand. 



