

20 BOTAMCAL GAZETTE [july 



5. The first marked internal differentiation in all three consists 

 of the gelatinization of a zone of hyphae in a region that will become 

 a part of the inner wall of the peridium. A zone of closely inter- 

 woven filaments just to the inside of this forms the boundary 

 between gleba and peridium. 



6. The origin and development of the peridioles is similar in 

 all. Each peridiole originates around a center, toward which the 

 ends of filaments converge. The structure of the walls of the 

 peridioles differs only in relative proportions. 



7. The first peridioles to be differentiated in Cyathits fascic- 

 ularis are toward the base of the gleba, and later other peridioles 

 develop above them. The peridioles at the base mature before 

 those nearer the top of the gleba. In C. striatus and Crucibulum 

 vulgar e the peridioles appear almost simultaneously throughout 

 the glebal region; but the upper peridioles in C. striatus mature 

 before the lower ones; while in C. vulgare the development is more 



uniform. 



form from 



m 



The 



similar in both. 



le most marked difference between Crucibulum and 



structure of the walls of the Deridia. In Cvathus 1 



middle 



pseudoparenchymatic layer is present which is entirely wanting 

 in Crucibulum. 



10. During the expansion of the basidiocarp in Cyathus the 

 peridium is pulled off from over the glebal region, leaving parts of 

 the ungelatinized ground tissue to form, for a time, a thin covering 

 (the epiphragm). In Crucibulum the epiphragm consists of the 

 undifferentiated primordial tissue covered with branched hairs. 

 This undergoes gelatinization at maturity. 



11. The spores of all are constantly binucleate. 



I wish to express my indebtedness to the late Professor George 

 F. Atkinson, who saw a first draft of this paper, for a number of 

 helpful suggestions, to Miss Gertrude E. Douglas for the 



collection of much of the material used in the study of Crucibulum 

 vulgare, and to Dr. E. A. Burt for going over the manuscript of 



