270 BOTANICAL GAZETTE [NOVEMBER 
clear space that once surrounded them on three sides, and at this 
point they are now in a subapical position, almost in contact with 
the veil tissue. Remnants of the clear space still remain on both 
sides, but it is rapidly being filled by the growth of fundamental 
tissue which has arisen from the mycelium, and which has pushed up 
before it the mass of generative hyphae. The intrusion of funda- 
mental tissue from below has its beginning at an earlier stage, and 
indications of it may be seen on both sides at the base of the asco- 
carp in fig. 4. Judging from these two stages, it would appear 
that the most rapid formation of new tissue, at least during the 
early stages, takes place at the base rather than at the apex of 
the ascocarp. 
The subapical position of the generative hyphae is retained as 
the ascocarp increases in size, until the apex broadens out to form 
the cap. These hyphae then pass into the cap, and there some of 
their branches are transformed into procarps. This transformation 
entails an increase in diameter, the assumption of an irregular 
coiling habit, and the development of a process which passes from 
the procarp more or less directly through the veil to the exterior. 
These processes are to be regarded as trichogynes. The history of 
the procarps will be discussed in greater detail subsequently. While 
at this stage, the upper portions of the cap are densely filled with 
generative hyphae, procarp complexes, and their proliferations the 
ascogenous hyphae, interwoven in an almost inextricable tangle 
g. 6 
It is important to note that until this time no sign of a hyme- 
nium has made its appearance. Very shortly after paraphysis for- 
mation commences, however, and following this the ascogenous 
hyphae proceed with the proliferation of asci. Paraphyses in 
Cudonia arise from undifferentiated vegetative elements. ‘They are 
multiseptate, in this respect differing from the description given for 
this species by Duran», and their cells are multinucleate. At first 
they are straight, but later become strongly recurved at the tips, 
which are also somewhat enlarged. 
The veil still completely invests the entire ascocarp, including 
the hymenium: Over the hymenium it has become separated from 
the tissue of the subhymenial region, with, indeed, no attachment 
at all except at the edges (fig. 7). Nevertheless it does not cease 
