1922] DUFF—GEOGLOSSACEAE 273 
TRICHOGLOSSUM HIRSUTUM 
The very young fructification of Trichoglossum hirsutum shown 
in fig. 22 measures 175 u in height, and is of simpler structure than 
the corresponding stages of either Cudonia or Spathularia. It is 
composed of compactly intertwining threads, the walls of which are 
comparatively thick and very dark. The hyphae all appear to be 
alike except for some, which, situated near the periphery and at 
right angles to it, extend beyond the ascocarp as straight, sharp 
pointed, unicellular hairs of greater diameter and thicker walls than 
the other threads. These setae correspond to similar ones described 
by Brown in Lachnea scutellata (5), and by Firzpatrick in Rhizina 
undulata (14). The setae of Trichoglossum are differentiated some- 
what earlier than in Rhizina, but do not contain a glutinous sub- 
stance such as is found in those of the latter form. The setae 
in Trichoglossum appear to be continuously produced over all 
peripheral parts of the ascocarp as it grows, but those that are 
earlier formed persist for some time, and come to be imbedded deeply 
in the new tissues that grow up around them. This is particu- 
larly evident in the hymenial region, where such growth is rapid 
(figs. 25, 26). 
Paraphysis formation commences reatively early at the apex 
of the ascocarp, before it has begun to broaden out to form the 
cap (figs. 24, 25). The paraphyses are large, unbranched, multi- 
septate threads, the cells of which are usually uninucleate, but 
apparently may come to contain several nuclei in exceptional cases. 
They are strongly coiled at the tips. 
Shortly after the paraphyses are organized, broadening of the 
apex of the ascocarp begins, and the hymenium extends over the 
entire surface of the globular cap as it takes shape and expands 
into the mature condition. 
From the figures it may be seen that the hymenium is of 
undoubtedly exogenous origin, and, moreover, the ascocarp itself 
is clearly gymnocarpous. Throughout the entire series of stages 
careful examination has failed to reveal the presence of an invest- 
ment of any kind. Trichoglossum hirsutum, therefore, adds one 
more species to those Helvellineae the hymenium of which is exposed 
from the first and the fructification gymnocarpous. 
