274 BOTANICAL GAZETTE [NOVEMBER 
The vegetative threads in a well developed ascocarp of Tricho- 
glossum hirsutum constitute a loosely interwoven tissue in which 
the individual cells of the threads are clearly distinguishable, even 
in thick sections. The threads pass with more or less uniformity 
from the base to the apex of the ascocarp. Since they are thick 
walled and consist of uninucleate cells, it is not difficult to see that 
hyphae of any other type are absent. The paraphyses make their 
appearance at an early stage (fig. 24), but ascus formation does not 
commence until the cap is well shaped. Then, just beneath the 
bases of the paraphyses in the subhymenium, there appear thin 
walled hyphae which, unlike the vegetative threads, form a very 
densely interlaced tissue. These constitute the ascogenous system 
of Trichoglossum. It is certain that no bodies such as the generative 
hyphae and procarps of Cudonia or Spathularia stand out as specific 
organs, nor is it possible to distinguish the homologues of procarps 
as distinct from the ascogenous hyphae. 
The ascogenous hyphae are short, since they are differentiated 
in such close proximity to the hymenium. Asci arise from them 
in the usual manner. The formation of an ascus is preceded by 
the inversion of the tip of an ascogenous hypha to form a crozier, 
the ascus growing out of the penultimate cell. The asci are very 
large, with dense protoplasm, and, especially after the dark multi- 
septate spores are mature, form very conspicuous objects in the 
hymenium (fig. 26). 
LEOTIA LUBRICA 
Since material of this form had been collected, and since pre- 
vious investigators are not in complete agreement on two important 
points in the life history, it was thought advisable to examine the 
material with a view to the possibility of shedding some further 
light on these points. The questions to which reference is made 
are those of the presence of the veil and the origin of the ascogenous 
hyphae, and attention was confined to these questions alone. 
Two specimens at approximately the same well advanced stage 
showed irregular tissue fragments overlying parts of the ascocarp 
and the hymenium (figs. 27, 28). These specimens were not the 
youngest of the fruit bodies examined. In no others, however, 
whether younger or older than these, were any similar evidences 
