278 BOTANICAL GAZETTE [NOVEMBER 
mycelium certain threads which have been called generative hyphae, 
and which are the most prominent central components of the simply 
organized fructification. These threads are clearly differentiated 
in the stained preparation from the rest of the ascocarp. So 
vigorously do they react to stains that all details of cell structure 
are obscured, even when nuclear stains are applied. 
The generative hyphae grow upward as the ascocarp increases 
in length, remaining subapically situated. When the upper part 
of the fruit body is differentiated into the cap these hyphae make 
their way into it, and in that position certain branches of this system 
appear that are enlarged, irregularly coiled, and deeply staining 
(fig. 6). These are the procarps. The procarp coils are continued 
upward by processes which can only be regarded as trichogynes 
(figs. 6, 8, 9). These are multiseptate and follow a more or less 
direct course toward the exterior. Where the procarps lie in a 
suitable position the trichogynes may be followed for their whole 
length, and may be seen to penetrate the veil tissue and to reach the 
exterior. Sometimes the procarps are deeply imbedded, however, 
and in these cases it is impossible to follow the trichogynes as far 
as the surface of the ascocarp. 
The number of procarps formed in this manner in a single fruit 
body appears to be very large, although it is not possible to count 
them, on account of the way in which they are interlaced and 
because of the presence of masses of generative and ascogenous 
hyphae. Some idea of their number may be conveyed, however, 
when it is known that as many as seven distinct trichogynes have 
been counted in a single longitudinal section 10 u thick through a 
region of the cap in which they are numerous. Their distribution 
is irregular. Some of them are formed closely under, or even partly 
imbedded in the veil tissue, while others are situated at a distance 
from the surface in the direction of the center of the cap (figs. 6, 
8, 9). Sometimes they are localized in one or two restricted por- 
tions of the cap, being entirely absent from others. 
Fig. 11 is a portion of an unusually loosely organized procarp. 
This section does not contain any of the upper portion of the pro- 
carp, but it shows most of the lower part and the generative 
hypha from which it arose. The latter follows a somewhat 
