1922] DUFF—GEOGLOSSACEAE 281 
SPATHULARIA VELUTIPES 
The origin of the generative hyphae in Spathularia has already 
been described. Mention has also been made of the fact that they 
come to occupy such a position in the ascocarp that when the 
hymenium has been formed they will lie just beneath it. Apart 
from their somewhat larger size, conspicuous staining qualities, and 
restricted position in the ascocarp, the generative threads do not 
appear to differ from the vegetative hyphae. The cells of both 
contain from one to several nuclei. 
After the paraphyses have been formed, procarps grow out from 
some of the cells of the generative hyphae. The procarps are of 
relatively large proportions and become ver ¥y str ongl y I i! hil 
They are curious conspicuous complexes of hyphae, Attention 
has already been called to them, and some of them are illustrated 
in figs. 19-21 and 43-45. They appear to exhibit no uniformity of 
structure, but their cells are all multinucleate, and frequently dis- 
tinct pairing of nuclei is observable (figs. 43-45.) 
Ascogenous hyphae arise from these procarps. This is very 
easily demonstrated where the hyphal complex lies particularly close 
beneath the hymenium, and where the ascogenous hyphae pass 
directly into the hymenium. Under these circumstances the ascoge- 
nous hyphae take on something of the staining qualities of the cells 
from which they arise, and they may be traced with ease into the 
hymenium, and there may be seen in connection with asci. Fig. 21 
is a plexus of such ascogenous hyphae arising from a group of large 
cells, some of which are visible. This large group may be traced 
through a series of sections 7.5 u in thickness. The origin and 
termination of these threads are not visible in all sections, but by 
following through the series their connections in both directions are 
easily ascertained. 
Fig. 19 affords another illustration of this type, showing two 
enlarged procarp cells, somewhat depleted of their contents, one 
containing a single nucleus and the other two nuclei. From these 
cells deeply staining ascogenous hyphae are to be seen passing out- 
ward, following a very irregular course to the hymenium, and there 
in direct union with young asci. The group of uninucleate asci of 
about the same age shown in this figure appears to have arisen 
