1922] DUFF—GEOGLOSSACEAE 283 
On the other hand, Brown (4) has figured the remains of what 
he interpreted to be an ascogonium, a structure found by him in one 
case only, at the base of an older ascocarp than the youngest with 
which Dirrricu presumably worked. The ascogenous hyphae pass 
up from the ascogonium, and may be traced to the cap, where they 
become much branched, finally finding their way to the hymenium. 
The writer’s material did not include stages young enough to make 
possible a direct confirmation of either of these statements, but it 
did show hyphal complexes just previous to the formation of the 
first asci strikingly suggestive of those already described for 
‘ Spathularia. 
At this period the ascogenous hyphae appear under the hyme- 
nium as deeply staining threads in various stages of crozier formation 
and ascus proliferation, and since they are not numerous as yet, they 
are easily distinguished and recognized. Beneath them in the cap 
may be seen at the same time very conspicuous groups of greatly 
enlarged cells. Some of these enlarged cells are quite empty, some 
contain light vacuolated protoplasm, and others are densely filled 
with deeply staining contents, identical in appearance with those 
of the ascogenous hyphae. These enlarged cells are usually 
rounded, occurring singly or in groups, but they may assume any 
of a great variety of forms. Their distribution in the ascocarp 
follows no regular arrangement whatever. They contain from one 
to several nuclei which are variable in size and sometimes may be 
very large (figs. 47-53). From these large cells ascogenous hyphae 
arise. This is readily demonstrated wherever they occur close to 
the hymenial region. 
Brown has described what he regards as storage cells in the 
ascocarp of Leotia. Of these he says: “While the hymenium is 
being differentiated some of the vegetative hyphae give rise to 
large storage cells... . . These large storage cells are formed in 
rows and give rise to paraphyses. The storage cells are at first 
multinucleate, but the nuclei usually fuse as growth proceeds.” 
He affirms that the nuclei of these cells, where they are more than 
one, may be very unequal in size, and the fusion nuclei are some- 
times of extraordinarily irregular form. The writer has not been 
able to find storage cells which exactly correspond to those de- 
scribed by Brown. The only large unusual cells noted were those 
