1922] SPESSARD—LYCOPODIUM 4II 
those of fig. 112 are equally distributed over the fungus region of 
L. lucidulum. 
In spite of these observations, there is another which needs to 
be noted, and which may cast doubt upon them. The structures 
shown in fig. 124 were found on the epidermis of both kinds of 
prothallia, That shown in fig. 125 was never found except in the 
decayed portions of both kinds of prothallia. I suspect that the 
latter is only the maturer stage of the former. The ‘‘spores” of 
fig. 126 were drawn from a ruptured heavy walled structure shown 
in the lower lefthand corner of fig. 125. There were sixteen spores. 
All these facts point to the ascomycetous nature of the fungus, 
but since these last named structures were all found without the 
epidermis or within decayed regions where external fungi would 
have access, it is conceivable that they have no relation whatever 
to the endophytic fungus. The internal mycelium was so constantly 
segmented, however, that the Oomycete group has little chance 
of being recognized. 
While the evidence presented is in no respect conclusive, it 
throws some doubt upon the Pythium theory as to the identity of 
the endophytic fungus of Lycopodium prothallia. The writer dis- 
likes very much to leave so important a question unsettled, but 
it seemed best at least to record what had been observed. 
Summary 
1. The prothallia of two more species of Lycopodium have been 
discovered. 
2. The sex organs of L. lucidulum are primitive, being frequently 
mixed in nature. 
3. The prothallium of L. obscurum var. dendroideum shows a 
form transitional between the L. annotinum and L. complanatum 
spe 
. The sex organs of the last named species are invaded by the 
ihe fungus and consequently show deformities. The 
antheridia and rarely the archegonia occur in enormous masses. 
5. The sex organs develop individually as described for the 
genus, and acropetally in the two species. 
