1862.] TYLOR WEALDEN FOOTrEINTS. 249 



The peculiar interest of the plaster cast now exhibited b}^ Mr. C. 

 S. Mann, of Eltham, taken from one of the best impressions visible 

 on the beach, is, that it represents what I believe to be the foot- 

 print of probably the hind foot of an Iguanodon, standing upon a 

 ripple -marked surface of sandy mud sufficiently hard to retain an 

 exact impression. The pressure of the foot has raised the sand sur- 

 rounding the impression about half an inch above the ripple-mark, 

 at the same time turning over some shells of the genus Cyrena, 

 which may be seen in the disturbed mud. 



Professor Owen's figure of the bones of the foot of the Iguanodon, 

 above referred to, exhibits phalanges having similar proportions, 

 and similar relative position, to the impressions visible in the newly 

 discovered footprint from Hastings, which measures 24 inches from 

 the toe to the posterior margin of the cup-shaped depression which, 

 I suppose, marks the heel of the Iguanodon, and is 3 inches in dia- 

 meter and |, inch deep. 



The impressions made by each of the three toes are well defined : 

 the middle one measures 11 inches in total length, 6 inches at the 

 posterior margin, widening to 7 inches, and then tapering doAvn 

 to 2 inches at the depression of its anterior extremity, made by 

 the ungual phalanx, which probably penetrated deeply into the 

 mud. The impression of the dextral or exterior toe is 9 inches 

 in total length, and 6 inches wide, tapering to an obtuse point, 



I inch wide at the claw ; that of the sinistral or internal toe is 



II inches long and 5 inches wide, tapering to 1 inch, with an 

 irregular cup-shaped termination as in the right toe. 



The posterior margin of the impression left by the exterior toe 

 commences at a point very much posterior to that left by the central 

 toe; while the posterior margins of the central and internal toes are 

 more nearly level with each other. 



The animal appears to have been walking in a direction nearly at 

 right angles to the ripple-marks, turning his foot a little on one side, 

 so as to give a slightly oblique direction to the footprint. The 

 animal appears in this instance to have left an impression of a rest- 

 ing foot, which is much more distinct than, and also difters in cha- 

 racter from, others of the associated imprints, which were apparently 

 made by feet of an animal in continuous motion. 



These remains occur in the upper part of the East Clifi*, near the 

 junction of the shales (known as " Tilgate Beds ") and the Hastings 

 Sand proper, corresponding in position with the strata of other loca- 

 lities in which osseous remains of the Iguanodon have been found. 



If, as I am disposed to do, we may really regard these trifid, 

 pachydactylous, and apparently uniserial imprints and casts of im- 

 prints as the '^ spoors " of quadrupeds, and not of bipeds, and if we 

 refer them to the thick-footed, three-toed Iguanodon, we have indi- 

 cations of the tracks of that great reptile at several places and on, at 

 least, two horizons in the Wealden area. The footprints already 

 described by Beckles are, first, from grey sandy shales at and near 

 Couden and Eoxhill, west of Hastings. Here the tracks were repre- 

 sented by numerous imprints on the surface of the shales. The foot- 



