On the Gustatory Organs of some Edentata. 337 



The secondary papillae are fewer in number, there being not more 

 than five or six to a papilla as a rule, and resemble somewhat the 

 stout, hard spines of the Carnivora. Many of the papillae are sur- 

 moimted at each lateral border by a single recurved spine, the space 

 between being packed with epithelium. When viewed in horizontal 

 section they present a horseshoe-shaped cavity. These papillae may 

 be looked upon as representing an intermediate type between the 

 „coronate" and „fasciculate" papillae of Marsupialia and the corre- 

 sponding papillae of still higher forme. Another, and more simple form 

 of papilla occurs on tliis tongue near the lateral margins. It consists 

 of a single papillary upgrowth of the mucosa, overspreading which is 

 a layer of stratified epithelium. From the bed of epithelium rises a 

 single sharp-pointed spine. The spine is cornified at its upper part, 

 and directed inwards and backwards. 



Gustatory Structures. 



The Circumv aliate Papillae, — The papillae were not developed 

 alike in all the specimens. While some of them resemble the papillae 

 of higher animals, others approach more closely the marsupial type. 

 The former, or more recent type are 1,3 mm. in diameter and 1,1 mm. 

 in height. They are flattened on top, and barely reach the level of 

 the lingual surface. The bulbs are disposed around the lower part of 

 the lateral area in eighteen closely-packed tiers. The circumvallate 

 papillae of less recent type are taller than the foregoing, and their 

 sides converge as they approach the opening of the trench. Their 

 lateral area is filled with bulbs to within a short distance of the top, 

 there being often thirty tiers of them. They measure 0,051 mm. in 

 length and 0,030 mm. in breadth. Serous glands are quite abundant, 

 the ducts opening into the trenches at their base and sides. 



The Lateral Gustatory Organs. — The lateral gustatory organ 

 of Dasypus villosus in not unlike that of Procyon lotor (described by 

 the writer in the „Journal of Anatomy", vol. XXIV, 1890). The super- 

 ficial examination of this region showed several irregular slit-like open- 

 ings, but I only succeeded in obtaining sections through one of them. 

 These openings meas. 0,20 mm. in width, and lead into a large irregular- 



luteruatiünnle Monatsschrift fur Auat, u, Phys. VII. 22 



