THE GKNUS ROSA 



255 



-N^" 



Text -fig. I. a. 7i'. flexibilis ( Eiicanincv ). 

 b. R. Rcuteri (Afzeliamt). 

 To sliow difference in stigma lieads. 



second they are reflexed (rarely spreading) and decidiunis 

 (figs. 9, 12, 13, pi. xviii; figs. 14, 15, 16, 17, pi. xix; fig. 

 20, pi. XX ; figs. 26, 27, pi. xxi), falling before the fruit 

 ripens. Finally, the first section of any given pair ripens 

 its fruit markedly in advance of the second. It will thus 

 be apparent that one section-species of each pair of the 

 CanincB is fitted for self pollination and northern* latitudes, 

 and the second for insect pollination and more southern 

 climes, thereby indicating that the parallel evolution of the 

 allied pairs from the primitive Gz/////^ ancestor was possibly a 

 response to environmental factors. In this parallel move- 

 ment we discern the first hint of the workings of the unfolding 

 principle of Elmer's. 



But progress did not end here ; in these groups (as in others 

 not studied by me) in Asia, America and Europe, a gradual 

 advance has occurred with the evolution of a multitude of 

 species-types, each species-type marking a definite halting 

 point in the onward march. Once again, as these stages 

 reveal themselves, we are bound to admit that throughout the 

 sections there is complete agreement, for each species-type is 

 common to all ; a further unfolding of the latent powers of the 

 sections has manifested itself. Again the evolutionary 

 principle followed is strictly orthogenetic in its action. 



Next, in each section-species, every species-type breaks into 

 two microgenes, in one of which the foliage is of a full green 



* And in more southern stations for mountainous habitats, as Gelmi so 

 clearly explains in the case of Rosa glanca in the Trentino 



