318 



Systematic Paleontology 



Cycadospadix milleryensis from the Permian of Autun, Prance/ The 

 seed-bearing sporophylls of Eenault's interpretation would then be 

 morphologically the fertile leaflets of a single compound sporophyll. 

 Within the whorl of " stamens " of the Cycadeoidea " flower," which were 

 wilted or shed quite early, an ovoid or conical, ovulate cone was borne. 

 The orthotropous ovules were situated at the ends of long slender pedi- 

 cels, and the interstices were packed with sterile appendages (inter- 

 seminal scales) which, with their expanded tips formed a protective 

 envelope for the developing seeds, suggesting an angiospermous pericarp 



A B 



Fig. 8. — A. Radial longitudinal section of an ovulate strobilus of Cyca- 

 deoidea, somewhat reduced. (After Wieland.) 



B. Semi-diagrammatic longitudinal section of a bisporangiate strobilus of 

 Cycadeoidea, about one-fourth natural size. (After Wieland.) 



as shown in fig. 8 in actual section at A, and semi-diagrammatically at B. 

 The gigantic " stamens," if they may be called stamens, which is a 

 term of very doubtful propriety, have not been found unfolded, but as 

 the American material is immature Wieland rightly supposes that they 

 eventually opened out as shown in part in fig. 8, B. 



* Renault, Bassin Houill, d'Autun, 1896, fasc. 4, pt. 2, p. 329, pi. 73, figs. 1-7. 



