450 F. vox HUEXE CARNIVOROUS SAURISCHIA IX EUROPE 



Specimexs FPiOM Lower Lias axd Lower Oolite of Exglaxd 



In the English Lower Lias and in the Lower Oolite there are forms 

 which, while closely related to the genus Megalosaurus, yet do not really 

 belong to it, since they lie somewhere between Teratosaurus and Megalo- 

 saurus. For the present I shall not assign a name to this genus. The 

 tibia of these specimens is characterized by a rudimentary crista lateralis 

 at the lateral side of the proximal end and below its head. In Terato- 

 saurus this crest is still missing and in Megalosaurus it is strongly de- 

 veloped. The species from the Parker collection (Oxford) from Xether- 

 comb (Humphriesi horizon of the Lower Oolite) I have named ''Megalo- 

 saurus" netherconihensis n. sp. Its pubis is rodlike in the distal 

 (anterior) portion. The teeth are similar to those of the true Megalo- 

 saurus hucklandi, but, among other differences, are somewhat thicker 

 than in that species. 



The true Megalosaurus hucklandi occurs only in the Stonesfield slate 

 just below the Great Oolitic. So much of the skull is j^reserved that a 

 restoration, within certain limits of error, is possible. This skull ex- 

 hibits some resemblance to Antrodemus. The cervical vertebrte are high, 

 relatively short, and opisthocoelous. ]^either the cervical nor the dorsal 

 vertebrae show the j)leurocentral excavations so sharply circumscribed or 

 so deep as in Streptospondylus or in Antrodemus. The number of pre- 

 sacral vertebra? is not known, but there is no valid reason why it should 

 differ from that in Streptospondylus cuvieri, which has nine cervical and 

 fourteen dorsal vertebrae. The dorsal vertebra? show only pleurocentral 

 depressions, but not cavities. Below the diapophysis are strong support- 

 ing buttresses, with deep niches between them. The neural process is 

 broad and thick, but not very high. The sacrum, as is well known, con- 

 sists of five vertebrae. The anterior caudal vertebra? are as long as the 

 dorsals. The h^mapophyses are like those in Antrodemus. The scapula 

 is long, slender, and straight, as in Antrodemus, and its processus del- 

 toideus is very high. The humerus, which is robust, is about half the 

 length of the femur and a little more than half the length of the scapula. 

 The ulna and radius are scarcely more than half as long as the humerus 

 and are extraordinarily stout. Unfortunately the manus is not known. 

 In the pelvis the pubis is a narrow and distally rodlike bone, but is not 

 completely known. The proximal expansion containing the ol)turator 

 foramen disappears within a short distance of the j^roximal end. The 

 ischium exhibits an angle in the middle of the shaft. The ilium is rather 

 large and has a broad anterior spine. The trochanter major on the femur 

 is a broad fanlike crest for the ilio femoral muscle, corresponding there- 



