﻿Yol. 64.] LOWER SAI^DSTONES OF THE EXETER DISTRICT. 499 



which fall slightly in front of those of the pes, are comparatively 

 faint, each digit being represented by a small round dot, as if only 

 the tips of the digits had touched the ground, and then quite 

 lightly. The pes, on the contrary, has sunk deeply in, and the 

 marks of each toe are elongated ovals — somewhat sharper in front. 

 Evidently the centre of gravity of the animal must have been 

 situated in the pelvic region. 



Both limbs had four digits, the one farthest from the median line 

 being smaller than the other three. If this may be regarded as 

 the fifth digit, then 4 was the longest, 3 slightly shorter, and 

 2 again slightly shorter. Some of the prints only show the marks 

 of the three longest digits. 



Although there is so much difference in the weight carried by 

 them, the manus seems to have been of much the same size as 

 the pes. 



There is no trace on any of the slabs of a tail having dragged 

 upon the sand ; but it is difficult to understand how the centre of 

 gravity can have been so far back, unless the head and fore part 

 of the body had been counterpoised by some such appendage. 



The longest and much the most perfect track is that which I 

 have called E. This runs along the length of a large stone which 

 Messrs. Collard & Sons most readily cut from a much larger slab. 

 The track (which is a cast) is about 1'5 metres long, and contains 

 thirty pairs of prints, many of which are quite sharply defined. 



The animal had all the characters of the last described, exce^Dt 

 that the digits 5 and 2 were nearly equal, and that the creature 

 was somewhat smaller. Its full stride was 9 centimetres, and the 

 centres of manus and pes were only 2 cm. out of the median line. 



The print of the manus generally falls about 1 cm. in front of 

 that of the pes, but the spacing is irregular, and towards the end 

 of the track the animal evidently paused for a moment, as if 

 hesitating; There is no indication of the small digit having diverged 

 from the others, but 4, 3, and 2 do diverge slightly inwards towards 

 the median line. 



The track shows that, as the animal walked, it sometimes threw 

 nearly all its weight on the right limbs, sometimes on the left. 

 This cannot be an illusion due to partial obliteration, nor can it be 

 due to variations in the texture of the sand, because the prints are 

 uniformly sharp. In two cases the left manus has only left a slight 

 mark of the two longest toes, and in one case it has made no mark 

 at all. The corresponding marks of the right limbs are particularly 

 deep. Possibly the centre of gravity may have swayed from side to 

 side with the movements of the head or tail. 



In no case is there anything to suggest either claws, a sole to 

 the foot, or a fifth digit. The prints do not resemble any of those 

 figured in the reports of the British Association Committee now at 

 work on the Elora and Eauna of the Trias, nor any others at once 

 accessible. They are least unlike the toe-marks of some animals 

 which are classed as Cheirotheroid, but differ from them in the 

 absence of a divergent digit, the absence of anything like the palm 



