1894.] NEW GENERA OF EARTHWORMS. 383 



two couples of each side, while the ventral couple of one side is 

 separated from its fellow of the other side by a distance of 2*5 mm. 

 I could not find any trace of setae at all upon the first five 

 segments of the body. If this absence of setae upon the head end 

 be confirmed it is of interest, as this cephalization is rare among 

 the Cryptodrilidae, though a common character in the family Greo- 

 scolieidse. Geodrilus in fact is the only Cryptodrilid in which I can 

 recall anything of the kind. Segments vii.-xii. are bi-annulate. 

 The dorsal pores are very obvious. They commence on the border- 

 line of segments x./xi., possibly one or two segments earlier. 

 There are three of these pores upon the clitellum — one marks its 

 posterior boundary, while two He on the first two segments. The 

 clitellum is rather extensive, occupying segments xiii.-xxii. The 

 median ventral region behind the male pores seems to be free, at 

 any rate to a large extent, of glandular tissue. The two male 

 pores lie upon segment xvii. ; they are highly conspicuous and are 

 transversely elongated orifices, which correspond in position to the 

 missing ventral setae of the segment. Neither the oviducal nor the 

 spermathecal pores were visible. The body-wall of both the present 

 species and Millsonia nigra is exceedingly tough. Mr. Millson 

 informs me that this was also the case during life. 



Vascular system. — The dorsal blood-vessel of the worm is single 

 from end to end of the body. Iu segments xvi. and xvii. it is 

 distinctly dilated, forming thus a kind of heart. A local dilata- 

 tion of the dorsal vessel is not unknown, though rare, among the 

 Oligochaeta. In the Geoscolecid Microchceta I and Benham have 

 described the same kind of thing, while many Enchytraeids also 

 show a dilatation of the dorsal blood-vessel just after its emergence 

 from the peri-intestinal sinus (or plexus). I regard all these local 

 expansions of the dorsal blood-vessel as having some relation to the 

 heart of the Arthropods. The last pair of circumcesophageal 

 trunks are in segment xii. ; the five pairs which lie in front of 

 these are equally large. 



Intersegmental Septa. — The first distinguishable septum lies 

 between segments iv./v. It is tolerably stout and runs in a straight 

 course across the body. The four following septa are excessively 

 delicate and are pushed back by the stout gizzards so as to have 

 lost their definite relation to the segments which they separate. 

 After these thin septa come a number which are very strong 

 and muscular. The septa dividing segments ix./xvii. are stout, 

 diminishing in thickness posteriorly. The anterior of these and 

 those which lie in front of them as far back as septum xiv./xv. are 

 traversed by or give rise to muscular straps which are also 

 attached to the parietes and to the alimentary canal. 



Nephridia. — I do not give a long account of the nephridia under 

 the present species as they are constructed upon the same plan as 

 those of Millsonia nigra, in which species it so happens that I inves- 

 tigated them more closely. The peculiarity of the nephridia of this 

 genus, to which I have already referred, is not quite so strongly 

 marked in the present species as it is in the next to be described. 



