Vol. 68.] THE ' PERMIAN ' EOCKS OF HAMSTEAD. 673 



with those of this Hamstead type Hg occur commonly in the 

 Middle and Upper Rothliegende of Thuringia and in the Roth- 

 liegende of Moravia, Bohemia, and Silesia. The Hamstead speci- 

 mens of Hg have furnished no measurable fore footprints, hut the 

 hind footprints show a very close resemblance to those described 

 by Prof. Pabst as Ichnium gampsodactylum. The sole difference 

 between the Hamstead and the German forms is the more frequent 

 absence in H5 of the print of the first toe, which is only shown on 

 one specimen, and then but 0*7 cm. long; whereas the average 

 length of this toe-print in the German I. gampsodactylum is 1-3 cm. 

 It must be noted that there is no indication of this toe, on two 

 out of the eight measured German specimens. The description 

 of H5 given on p. 670 accords in all respects with that given by 

 Prof. Pabst of /. gampsodactylum. 



Sub-Type Hs^. Ichnium gampsodactylum (minus) Pabst. 



The footprints in type Hs^ (fig. 26, p. 672) appear to agree almost 

 exactly with the small footprints of /. gampsodactylum separated 

 by Prof. Pabst to form the subspecies minus (fig. 27). In type H5 

 no fore footprint was clearly defined, but in H5„ both hind foot 

 and fore foot are well shown, and in both the resemblance to the 

 German form is very plain. It is to be noted, however, that the 

 first toe of the fore foot and the fifth toe of the hind foot are absent 

 from the Hamstead impressions, although present on some of the 

 German specimens. The Continental specimens are derived from 

 the Middle and Upper Rothliegende of Thuringia, and from the 

 Rothliegende of Silesia, Bohemia, and Moravia. 



Dr. G. Hickling ^ figures and describes a footprint from Penrith, 

 collected by Mr. G. Varty Smith, which resembles closely the 

 Hamstead type H5 {Ichnium gampsodactylum). 



Type He. Compare Ichnium aceodactylum (minus) Pabst. Two 

 specimens (casts). (See fig. 28, p. 674.) 



The specimens show tiny footprints, which are longer than they 

 are broad- The ball of the foot is long and has a distinct heel. 

 Five toes are indicated, which are long and taper gradually to a 

 point. Each is straight throughout. The fifth toe diverges most 

 from the line of the other four, which run closely side by side. 

 The first is the shortest, and the third and fourth are the longest. 

 In the connected track shown on one of the specimens, the fore and 

 hind feet are not clearly distinguishable, owing to their close 

 approximation in size and to the fact that track-measurement I 

 is practically equal to track-measurement II. The gait is not 

 alternating, but opposite. 



^ 'British Permian Footprints' Proc. Manch. Lit. & Phil. Soc. vol. liii 

 (1909) no. 22, p. ] . 



Q. J. G. S. No. 272. 3 b 



