THE GENUS PUMARIA IN BRITAIN 21 



racemes, and very obtuse fruits. But the forms connected with 

 these two types are legion, and some of the Madeira plants that I 

 have named F. muralis var. Loiuei seem to supply a link with 

 this Cornish plant, especially in respect of the long racemes and 

 the form of the corolla. I therefore think that this new plant is 

 best placed as a new subspecies neglecta of the aggregate 

 F. muralis. 



It is no doubt extraordinary that another new fumitory should 

 thus be found in Cornwall, on this occasion, in an isolated field. 

 I think, though, that it was almost certainly introduced at some 

 perhaps remote date from the Continent, and possibly from Spain 

 with the new species next to be described, although I have seen 

 no similar Spanish example. Much more work, I quite believe, 

 has yet to be done in the South of Europe with the Fumarias of 

 this polymorphic subsection, and probably at some future date 

 the headquarters of this particular form will be discovered. The 

 only specimen at the British Museum or at Kew at all re- 

 sembling F. neglecta is one at the latter place, very questionably 

 labelled *• M. Ital. Exsicc, A. Fiori, A. Beguinot, R. Pampanini, 

 Fumaria serotina, Guss, var. coiifusa, Jord. Bordighera, C. 

 Bicknell, Mar. 1904," which I believe to be a shade-form of 

 F. Gussonei. 



Specimens of this new plant were sent unnamed to the 

 Watson Exchange Club by Mr. F. H. Davey in the winter of 

 1907-8. 



The characters of the plants referable to F. muralis, F. Borcei, 

 and F. neglecta are so divergent that it may be held inconsistent 

 to unite them under one species while other fumitories, apparently 

 differing no more widely, are considered specifically distinct. But 

 it can hardly be questioned that, either in Britain or abroad, 

 nearly all of these muralis forms pass into one another, which is 

 not known to happen in the case of our other species. It may 

 be recalled, too, that the comparatively large fruits so pre- 

 valent in the British forms are also characteristic of the 

 common British variety of F. capreolata and may, with all of 

 these plants, be a result of the moist climatic conditions of our 

 islands. 



The features common to all the muralis forms, besides the 

 nearly or quite smooth fruits, are a foliage approximating to that 

 of F. capreolata : ovate, peltate sepals, toothed chiefly towards 

 the base ; a pink corolla ; and a dorsally compressed tube of the 

 upper petal, with a distinctly spathulate dilation of the invariably 

 dark purple wings. 



From this aggregate species F. Eeuteri Boiss. and F. Munbyi 

 Boiss. are clearly separable by their narrower leaf-segments and 

 smaller, compressed fruits, while the little-known plant, F. senium 

 Boiss., is apparently quite distinct, owing to its narrow lanceolate 

 sepals. F. gaditana Haussk., of the subsection Capreolatce, may 

 be distinguished by its much longer peduncles, more entire sepals, 

 and paler flowers. 



