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1883, ] 201 [Packara, 
from the so-called under lip of Chilognaths ; they are not united, and are 
separate, cylindrical, fleshy, 5-jointed appendages, but as Newport states 
“connected transversely at their base with a pair of soft appendages (c, ¢), 
that are situated between them, and which, as I have already stated, I re- 
gard as the proper lingua, as they form the floor of the entrance to the 
pharynx.’ These 5-jointed appendages are Mr. Newport’s ‘‘ maxillary 
palpi;’’ his true maxille being the homologues of the ‘‘mandibles”’ of 
Chilognaths, 
The portion of the head of Scolopendra and other Chilopods, thus far 
considered, together with the antenne and proto and deutomale, we con- 
sider as homologous with the entire head of Chilognaths ; the basilar seg- 
ment of Newport, and the two pairs of head-appendages have no homo- 
logues in the head of Chilognaths. They are rather analogous to the 
maxillipedes of Crustacea, and nothing like them, speaking morphologi- 
sully, exist in other Tracheata. We therefore propose the term malipedes 
(mala, jaw ; pes, foot, or jaw-feet) for the fourth and fifth pair of cephalie 
appendages of Chilopoda. At the same time it is easy to see that they are 
modified feet; especially when we examine the last pair in Scolopendra, 
which are attached to a true sternite, and see that they are directly homo- 
logous with the feet and sternite of the same animal. 
The first pair of malipedes are the ‘labium and palpi’? of Newport ; 
the ‘first, auxiliary lip’’ of Savigny. They, however, bear little resem- 
blance to an insect’s labium and labial palpi. They are separate, not coa- 
lescing in the middle as in the labium of Hexapods. The so-called labial 
palpi are 4-jointed, with an accessory plate. They arise directly in front 
of the ‘basilar segment’’ of Newport, but appear to have in adult life no 
tergite of their own.* 
The second pair of malipedes or last pair of mouth-appendages, are the 
poison fangs ; they are the ‘second auxillary lip” of Savigny ; the ‘‘man- 
dibles or foot-jaws’’ of Newport and subsequent authors. The dorsal plate, 
or what may be called the second malipedal tergite is the ‘basilar and sub- 
basilar plate’? of Newport. 
As to the number of segments in the head of Chilognaths, both mor- 
phology and embryology prove that there are but three ; in the Chilopoda 
five, Newport’s observation on the young recently hatched Geophilus 
(his Pl. xxxiii, fig. 3), shows that the sub-basilar plate is the tergum or 
scute of the fifth segment; and the basilar plate is consequently the 
tergum of the fourth segment, or second malipedal segment. The ster- 
nite of the sub-basilar plate is usually a very large plate, deeply in- 
dented in front in the middle, with teeth on each side, and forms the ‘‘la- 
bium’”’ of Newport. It may for convenience in descriptive zoology be 
termed the ‘‘pseudolabium.’’ 
> 
* Balfour also states, as we find after writing the above, that the basilar plate 
is really the segment of the poison claws, and may fuse more or less completely 
with the segment in front and behind it, and the latter is sometimes without a 
pair of appendages (Lithobius, Scutigera) Comp. Embryology, i, p. 225, 
PROC, AMER. PHILOS. SOC, XXI. 114. Z. PRINTED SEPTEMBER 17, 1883. 
