48 Memoirs of the Indian Museum. [Vor. VI, 
The salivary glands are small; they originate from the junction of pharynx and 
oesophagus as diverticula which are prolonged backwards each as a solid club-shaped 
mass of cells (fig. 3, sal.), the posterior end being rather broader than the stalk, and 
appearing to be somewhat curled inwards behind the pharynx towards the middle line. 
These structures are limited behind by the sheet of muscle-fibres passing downwards 
and backwards from the pharynx. In their club-like form and small extent of the 
lumen they resemble the structures described in Enchytraeus barkudensis (23). 
Septal glands are present in segments iv, v and vi; they are bulky, filling the 
segments at the sides of the oesophagus ; those in v and vi have forwardly extending 
ventral lobes in addition. ; 
Chloragogen cells begin in v, and are numerous in vi and behind. The oesophagus 
passes into the intestine without the appearance of any marked difference between 
these two portions of the canal. 
The chyle cells which are a character of the genus extend in this species through 
segments xiv—xviii. In these segments they form a considerable portion of the ali- 
mentary epithelium, indeed in a number of sections apparently the whole, though 
usually other more cubical cells are present along with them. ‘The chyle cells are 
elongated columnar cells, their peculiarity, as is well known, being the possession of 
a central canal which opens into the lumen of the gut. In the present species the 
canal is straight for the greater part of its length, but at the basal end of the cell it is 
often bent to a right angle or curved into a semicircle; the portion of the canal near 
its mouth seems to be ciliated, but only that portion. The free surfaces of the cells 
are ciliated ; the nucleus, containing a central karyosome, is situated at about halfthe 
height of the cell or a little lower. The lumen of the canal within the cell is bordered 
by a zone of hyaline non-granular protoplasm, and beyond this again is a zone of 
deeply-staining (with iron haematoxylin) granular particles. The cells are not in close 
apposition throughout their length; there appear to be considerable intervals between 
them ; the intestinal blood-sinus bathes their basal ends. 
The dorsal vessel can be traced backwards in sections as far, at least, as segment 
xv; in the posterior part of ix, just in front of its passage through the septum, a 
number of cells are aggregated inside its lumen, but these are not definite enough to 
be called a cardiac body. A sinus-like space containing blood can be traced in the 
intestinal wall as far forwards as segment vi. 
The nephridia begin from segment vii. The anteseptal portion is of relatively 
considerable size, perhaps a third as long as the postseptal, and the tube undergoes 
some windings in the anteseptal portion before piercing the septum, The duct appears 
to leave the lower surface at the posterior end. The determination of the characters 
of the nephridia in preserved material is not always satisfactory. 
The cerebral ganglion is rounded behind, concave anteriorly ; it is a little longer 
than broad, and lies above the buccal cavity, in front of the hemispherical pharynx. 
The ventral nerve cord shows a giant fibre dorsal in position. 
The festes were doubtfully identified ventrally in segment xi. The funnel begins 
anteriorly as a collar of cubical cells; the main bulk of the funnel, which succeeds the 
