AN ACCOUNT OF THE ORIENTAL PASSALIDAE (COLEOPTERA), 



BASED PRIMARILY ON THE COLLECTION IN THE INDIAN MUSEUM 



By F. H. GrA-VJoLvY, M.Sc, Assistant Supsrlntßndent, Iniian Museum. 

 (Plates XI— XIII.) 



CONTENTS. 



Page 



1. Introduction . . . . . . . . . . . . . . 177 



2. External anatomy with. sp2cial reference to taxonomic values . . . . 179 



3. Classification of Indo-Australian Passalidae . , . . . . . . 191 



4. Catalogue of specimens in the collection of the Indian Museum, with notes on those 



in certain other collections . . . . . . . . . . 204 



5. The variable species . . . . . . . . . . . . 262 



6. Geographical distribution and synonymy . . . . . . . . 265 



7. Appendix I. A revised classification of the Aceraünae . . . . . . 316 



8. ,, II. Keys for the determination of species . . . . . . 318 



9. ,, III. The genus Ifl/'g^miws and a remarkable new genus from New Guinea . 326 



10. ,, IV. Supplementary catalogue of specimens in the Indian Museum collection 330 



11. Summary and conclusions .. .. .. .. .. .. 336 



12. Bibliography . . . . . . . . . . . . . . 340 



13. Index .. .. .. .. .. .. ..345 



I. INTRODUCTION. 



The present Memoir is in effect Part II of the '' iVnnotated lyist of the Asiatic 

 Beetles in the collection of the Indian Museum." Its scope is, however, much larger 

 than that of Part I ; on this account and for other reasons which need not be detailed, 

 it has been decided to discontinue the publication of the " lyist " as a separate series. 



After I had commenced the catalogue of our Asiatic Passalidae it soon became 

 evident, on account of the present confused state of knowledge of the family, that 

 the value to specialists of the locality-records it was to contain would be greatly in- 

 creased by the inclusion of full illustrated descriptions of all the species referred 

 to ; and that if these were included its value to collectors in the E)ast would be still 

 further increased by the inclusion of a key for the identification of all genera known 

 from the Oriental Region. 



This led to an enquiry into the general principles on which the classification of 

 the family has been based; and to the conclusion that, by a little modification of 

 Kuwert's system, the Passalid fauna of the whole Indo-Australian area, together 

 with that of China and Japan, could be shown to be much more homogeneous and dis- 



