50 Selmar Schonland, [May 28,. 



development proceeds on the same lines as in belliera. Apart from 

 the unimportant differences as regards position and shape of the 

 entrance to the " indusium," Ave have to take also into account another 

 difference, which I will briefly discuss. 



In Selliera and probably in many other (if not in all) Goodeniaceae 

 the pollen is deposited in the " indusium " after having been dis- 

 charged by the anthers. But shortly afterwards two stigmatic lobes 

 begin to be developed, pushing the pollen out'of the cup, whence it is 

 carried away by insects in very much the same way as in Lobelia. 

 The stigma does not become receptive until it has grown through the 

 entrance of the cup ; in fact, the latter has the same biological function 

 as the tube formed by the anthers in Lobelia. Now in Cyphia 

 volubilis no stigmatic lobes are developed, and it is the inside of the 

 " indusium," which performs the function of a stigma. But even this 

 difference can, 1 believe, not shake the conclusion that what I have 

 now called the " indusium " in Cyphia volubilis is homologous to 

 what we are accustomed to call the "indusium" in Goodeniaceae.. 

 But to satisfy even the most scrupulous morphologist one might 

 perhaps say that the structure enclosing the " stigmatic cavity " is 

 homologous with the combined stigma and indusium of Goodeniaceae. 



It would be interesting to know for certain whether the other 

 species of Cyphia also possess a similar "indusium," although one 

 may conclude from Harvey and Sonder's description that this is the 

 case, and if found in all of them this character might be used as a test 

 whether the American genera Nemacladus, Parishella and Cypho- 

 carpus are so nearly allied to Cyphia, as is frequently supposed, 

 because the structure of the flower is somewhat similar in all of them. 

 I have, however, already expressed my doubts whether these 4 

 genera really form a natural group ( 1 ). 



Summary. 



1. The anatomical structure of the stem of Cyphia volubilis is 

 broadly speaking the same as in Lobelioideae and Campanuloideae, 

 The tuber contains Inuline as reserve-material. 



2. The flowers are not resupinated. 



3. The style contains a cavity in its upper portion, which com- 

 municates laterally with the open air by a narrow channel. The 

 tissues surrounding it are homologous with the combined stigma and 

 " indusium " of Goodeniaceae. 



4. It is advisable to include the Goodeniaceae among Cam- 

 panulaceae. 



('). " Natiirliche Pflanzenfamilien,'' iv., 5 p. 61. 



