i 
308 WHITE—OMPHALOPHLOLIOS, A NEW LEPIDODENDROID TYPE. 
type. These bolsters have large roundish or ovate triangular bosses, on 
which are placed the leaf scars and certain other structures. The large 
bosses were probably highly prominent in the uncompressed stems and 
were presumably composed, like the other portions of the bolster, largely 
of soft tissue that has proved very susceptible to distortion and varia- 
tion under the conditions of compression. Their prominence and lack 
of support well accounts for their partial concealment beneath the foids 
of the adjacent portions of the bolsters in the flattened impressions, as 
well as for their displacement toward the sides of the bolsters in many 
cases. The degree of deformation of the bolsters in this trunk exceeds 
the greatest variations from pressure I have seen in the bolsters of Lepz- 
dodendron Veltheemianum Stb., or L. clypeatum Lx. 
Pressure in a direction probably nearly vertical to the large boss evi- 
dently produced the rounded impressions described and figured from 
the originals by Lesquereux as Lepidodendron cyclostigma. From the 
lower and more prominent part of the large flattened boss two nearly - 
parallel, obscurely defined, broad, rounded, perhaps subcortical, ridges 
pass upward across the boss, and apparently a little beyond it, then 
seem to unite in a horseshoe curve or rounded angle. Within the apex 
of this loop, and apparently a short distance above the boss, is situated 
a rarely visible punctiform trace. I am unable to state whether this 
long, obscure, vertical loop is closed at the base to form an ellipse, though 
it shghtly affects that appearance in the pits figured in plate 21, figure 2. 
It may proceed on either side from the lateral wings of the leaf scar at 
the base of the boss. There is nothing on the specimens before me to 
indicate an attachment of any vegetative organ along its surface or margin. 
Certain very important points as to the relations of the second or oval 
boss to the leaf scar remain to be determined. At present it is not defi- 
nitely ascertained whether the oval boss, which in a few instances appears 
to be closed at the base and barely contiguous with the transversely an- 
gular or deltoid cicatrice which I have called the leaf scar, is actually 
distinct from that “scar” or whether it is organically connected there- 
with. The analogies with the other Paleozoic Lycopodinex, especially 
some of the Sigillarioid types, would at first glance lead us to inquire as 
to whether this oval boss does not itself represent a part if not the whole 
of the foliar cicatrix. ‘The evidence in support of such a supposition lies 
largely in the presence of the generally clear, narrow, very low marginal 
rim of the boss and its naturally suggestive similarity to the form of the 
cicatrices in some of the Bothrodendra. Continuing the parallel with the 
Bothrodendroid or Sigillarioid scar, it appears that in this case the trace 
at the upper end in the central oval depression may be the vascular trace, 
while, by reversing the position of the specimen, the punctiform trace, 
