PHYLOBLATTA TRANSVBRSALIS. 117 



The cubitus remains above the middle line of the wing in the basal third, 

 curving inwards and flattening in the outer two-thirds of the wing, and giving off 

 seven inward branches, the second alone forking low down near the margin. 



The anal area is small, and crossed by six oblique anal veins, the first and third 

 of which fork. 



The interstitial neuration consists of a close-set series of straight cross-nervures 

 which unite laterally as they cross the wider areas. 



The pronotum in close apposition with this wing has its inner surface upper- 

 most. This shows a central shallow concave area, bounded by two lateral ridges, 

 which die away as they approach the hinder border. The general outline is 

 broadly circular. 



The second tegmen is much less exposed than the first, and has its apex buried 

 under the second pronotum. So far as its structure can be determined, it agrees 

 with the wing already described. 



The second pronotum has been crushed, the anterior border being turned round 

 upon the base of the wing, and partly broken away. The posterior margin is 

 almost straight, and that of the anterior well rounded. 



Traces of two hind-wings are shown, one a little in advance of the more nearly 

 complete wing, and partly underlying it, and the other underlying the costal and 

 radial areas of the second incomplete wing. The first hind-wing shows a portion 

 of the distal extremity, with a series of incomplete veins, probably belonging to 

 the radius, median, and cubitus; the remains of the second hind-wing probably 

 consist of the distal radial twigs only. 



Affinities. — The distinguishing features of the tegmina are essentially those of 

 Phyloblatta, unless we except the interstitial neuration. This is composed of 

 straight cross-nervures, which unite laterally across the wide areas between the 

 main stems of the radius, median, and cubitus, but not producing anywhere a 

 meshwork such as is usually seen in the Phyloblattids. 



Handlirsch doubts the presence of a cross-neuration in the genus Phyloblatta 

 (' Proc. U.S. National Museum,' vol. xxix, p. 731, 1906), but there seems no reason 

 why it should not be present in the more archaic members of the genus, as they 

 all have an Archimylacrid ancestry, in which a cross-neuration is a dominant 

 feature. 



The species is closely allied to a form figured as Gerablattina sp. by Brongniart 

 (' Insectes Fossiles des Temps Primaires,' pi. xlvi, fig. 7, 1893), and repeated by 

 Handlirsch ('Die Fossilen Insekten,' pi. xxx, fig. 35, p. 295, 1900) under the name 

 of Blattoidea, sp. Handlirsch regards Brongniart's specimen as belonging either to 

 the Spiloblattinidge or to the Archimylacridas. Unfortunately the interstitial 

 neuration is not shown. If, as in these specimens, Brongniart's species possesses a 

 cross-neuration, or one in which the nervures unite laterally, it must be referred 

 to Pkyloblatta. 



