PSEUDOFOUQUEA CAMBRENSIS. 73 



lost from its tip since it was measured by Allen, who gives the total length as 

 41 nun. The greatest width is 15 mm. 



The subcosta is widely separated from the outer or costal margin in the base of 

 the wing, and gradually approaches and unites with it beyond the middle. The 

 radius is parallel with the subcosta throughout its length, and gives origin to the 

 radial sector basally. The radial sector diverges from the radius over the whole 

 of its course. It now shows but one inwardly directed branch, which forks at the 

 broken edo-e of the wins:. Allen's figure indicates that two more branches were 

 given off, both being undivided. The radius and radial sector occupied almost the 

 whole of the apex of the wing. 



The median vein forks in the basal fourth, the outer branch again forking 

 before the middle of the wing is reached. The inner branch of the median diverges 

 almost in a straight line from the outer branch, and also forks, and bears an 

 accessory twig upon the fourth branch. A feeble accessory twig appears to be 

 given off near the middle of the wing, but dies out in the integument. The cubitus 



Fig. 22. — Pseudofouquea cambrensis (Allen): restoration of whole wing-, natural size. Lower Coal 

 Measures (top of the Four Foot Seam) ; Llanbradach Colliery, near Cardiff. Basal portion of 

 wing in Mus. Pract. Geol. (no. 7272). Impression of apical portion of wing in the Welsh National 

 Museum (no. 13,120). 



is a remarkable vein, unlike that of any other fossil wing (Comstock, ' Wings of 

 Insects,' 1918, p. 106). For nearly half its length it passes in a broad curve to 

 the inner margin, giving off a series of alternate twigs upon its outer and inner 

 sides, those of the inner being weaker than those of the outer side. The two 

 outer twigs are strongly developed, while those on the inner side of the cubitus, 

 four in number, are weaker and shorter. The feeble continuation of the main stem 

 reaches the margin between the two sets of branches. Four anal veins are 

 distinguishable. The inner two arise from a common base, the outer two not 

 uniting. This is unlike the condition in Fouquea, where the anal veins branch off 

 regularly from a single stem. 



The area lying between the first anal vein and the main stem of the cubitus is 

 very wide — much wider, indeed, than any other area. 



The interstitial neuration consists of weak cross-nervures, except between the 

 base of the first anal vein and the main stem of the cubitus, where it is irregularly 

 reticulate. 



Affinities. — The characters of the cubital and anal veins definitely remove the 



species from the genus Fouquea, and the cubitus, with its strong, anteriorly 



directed twigs, and its feebler inner series, is wholly unlike that of any other 



insect, and would alone suffice to justify the generic rank given by Handlirsch. 



10 



