NEWER PLIOCENE BEDS IN BRITAIN. 283 
the convolutions of the frontal lobe much like those of the fossil, 
the three longitudinal convolutions (a, 6, ¢ in figures) being distinctly 
traceable ; but the brain of G. subgutturosa has not the lateral com- 
pression of the fossil; and consequently, although the same convo- 
lutions are seen, they are proportionately wider. In addition to 
this the external features of the skull are different, G. subguiturosa 
haying the horn-cores more upright and more curved, whilst the pit 
on the pedicle is further back and shallower. 
Gazella Bennettv is the species most like the fossil in the form of 
the exterior of the skull, the greatest observable difference being in 
the shape of the frontal fossa, the inner margin of which is less 
distinct than it is in the fossil ; and the pit on the pedicle of the 
horn-core is shallower and placed quite at the back. Mr. R. Ly- 
dekker has kindly lent me a skull of G. Bennettic, with permission to 
bisect it and take a cast of the brain-cavity ; and I find that its brain 
(fig. 8) more closely resembles this Pliocene fossil than does that of 
any other species with which I am acquainted ; andindeed it requires 
a close examination to trace any difference between them. However, 
although the same three longitudinal convolutions (a, 6, ¢) are to be 
seen, it will be noticed that the sulcus between the convolutions 
6 and ¢ is continued backwards, so as to separate these two conyo- 
lutions at their back part ; while in the fossil (fig. 4) they are joined 
at this point. There are other slight differences in the convolutions 
of the orbital region*. I am also indebted to Mr. Lydekker for the 
opportunity of examining the skull and cast of the brain of the 
Tibetan Gazella picticaudata; but both these parts have fewer 
characters in common with the fossil than have been shown to exist 
in the species above noticed. 
It now remains to be seen whether the Norwich-Crag Antelope 
will agree with any of the known fossil forms; for although no 
species of Antelope has hitherto been recorded from British strata, yet 
more than forty fossil species have been named from other parts of 
the world. Only a few of these, however, show sufficient resem- 
blance to our fossil to render it necessary to call special attention to 
them; and some, having been established on parts of the skeleton 
other than the skull and horn-cores, cannot be compared with the 
parts of the British Antelope at present obtained. 
All the fossil Antelopes from the Siwalik Hills which have been 
made known to us are larger than this British example, the one 
which comes nearest to it, Antilope porrectecornis, being at least half 
as large again, and having the horn-cores directed more outwards, 
with their inner sides definitely concave from end to end, while the 
frontal fossa is pyriform and not triangular. 
* Tt was thought that possibly different individuals might vary as to the 
characters of the skull or brain, and Mr. W.T. Blanford has very kindly placed 
at my disposal the skulls of two additional specimens of the Indian Gazella 
Bennettii for examination. Having taken casts of the interiors, I find that 
in their internal as well as external characters they agree exactly with Mr. 
Lydekker’s specimen. 
+ A list of allthe known fossil species, with references, is given at page 285. 
