272 S. W. WILLISTON NEW GENERA OF PERMIAN VERTEBRATES 



the only known amphibian, save Cochleosaurus,^ with an epicondylar 

 foramen, has been referred to the Microsanria or Lepospondyli by Cope, 

 Broili, and Jaekel, as also by myself provisionally, but, as I pointed out 

 in a recent paper,^^ the mode of attachment of the ribs is quite unlike 

 that of the typical microsanrs and like that of the modern salamanders. 

 Doctor Moodie has, because of this character, erected the new order of 

 Diplocanlia for the genus, and I believe the name is valid, though the 

 rank of the group is open to dispute. CrossoteJos Case, a form with 

 holospondylous vertebrae, found associated with Diplocaulus at Orlando, 

 has distinctly double-headed rrbs, with the capitulum attached to a facet 

 just back of the front margin of the vertebral centrum, the tubercle to 

 the extremtiy of a short diapophysis. True microsaurs have single- 

 headed ribs attached intercentrally, with the chevrons presumably at- 

 ' tached also as in reptiles.- In both Diplocaiilns and CrossoteJos the 

 chevrons are quite like those of the true amphibians, exogenous processes 

 from the middle of the centrum. But I am quite confident that at least 

 two distinct groups have been associated among what are called Micro- 

 sanria, and that one of them, with single-headed, intercentral ribs and 

 intercentral chevrons, must be dissociated into a group more nearly allied, 

 possibly identical, with the reptilia in a wide sense, while the other will 

 remain among the amphibia. As I am not able at the present time to 

 decide which of these groups includes the type form of the order, it is 

 premature to attempt the solution of the problem by giving new names. 

 This thing, however, may be said with assurance, neithei Diplocaulus nor 

 Crossotelos can be united with such forms as Eosauravus copei Will. 

 (nom. nov., Isodectes punctulatus Cope in part, Isodectes punctulatus 

 Moodie), or Sauravus costei Thevenin. Crossotelos has numerous slender 

 ventral ribs, very much like those of Labidosaurus and Captorhinvs 

 (Pariotichus) , of which I have found abundant evidence in new material 

 from Texas. 



The humerus of T rimer orliaMs (plate 15, figure 6) departs markedly 

 from the ordinary temnospondyle type, as do also the femora, in which 

 the adductor ridge is represented by a mere line on the ventral side of 

 the femur quite as in the reptiles, and unlike all otlier known Texas 

 amphibians. It is also remarkable for the small size of the lateral pro- 

 cess, which is separated from the head by a considerable free space. This 

 process is divided for the attachment of the several muscles into two 

 rugosities opposite each other on the front and hinder surfaces. The 

 median process, usually feebly indicated in the amphibians, is clearly 



» Brolll : Paleontographlca, 111, p. 15, fig. 3a. 



"Transactions of the Kansas Academy of Science, vol. xxli, p, 122, 



