P.N. WYSE JACKSON 
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Synopsis. A systematic appraisal of the partially silicified Lower Carboniferous bryozoan fauna of County Fermanagh has 
demonstrated a rich and diverse bryozoan fauna of which the fenestrate portion has been largely described earlier by other authors. 
This paper describes the remaining cryptostome, trepostome, and cystoporate elements of the fauna, as well as a few previously 
ignored fenestrate taxa. 
24 species are described (9 cryptostome species; 6 fenestrate species; 6 trepostome species; and 3 cystoporate species) of which 
6 are new species and 2 new combinations. The new species are the arthrostylid cryptostomes Hexites paradoxus, Nematopora 
hibernica, and Pseudonematopora planatus, the rhomboporid cryptostomes Rhombopora cylindrica, and Rhombopora hexagona, 
and the trepostome Leioclema indentata. The new combinations are Clausotrypa ramosa (Owen), and Rhombocladia dichotoma 
(M‘Coy). For completeness brief descriptions are given of the following taxa, which have been described more fully elsewhere: 
Rhabdomeson progracile Wyse Jackson & Bancroft, Baculopora megastoma (M‘Coy), Diploporaria tenella Wyse Jackson, 
Thamniscus colei Wyse Jackson, and Fistulipora incrustans (Phillips, 1836). 
The genus Leioclema and the species Streblotrypa pectinata Owen, Diploporaria marginalis (Young & Young), Dyscritella 
miliaria (Nicholson), Tabulipora howsii (Nicholson), and Tabulipora minima Lee are reported from Ireland for the first time. The 
following genera are reported from the British Isles for the first time: Hexites, Pseudonematopora and Clausotrypa. Lectotypes 
are designated for Diploporaria marginalis (Young & Young), Ichthyorachis newenhami M‘Coy, Rhombocladia dichotoma 
(M‘Coy), Dyscritella miliaria (Nicholson), Tabulipora howsii (Nicholson), and Tabulipora minima Lee. Nomenclature problems 
for several species have been clarified. A tabular and dichotomous key is given for the complete fauna (including taxa described 
earlier by other authors). Patterns of silicification show that replacement of calcified bryozoan zoaria by silica was delayed. 
INTRODUCTION 
Bryozoans comprise a significant component of Lower Carbonifer- 
ous faunal assemblages in Ireland. However, they are often 
fragmentary in nature which has made them difficult to study. 
Nevertheless, Lower Carboniferous bryozoans have been the subject 
of research since the mid-1800s when M‘Coy (1844) described 
many species. In the last thirty years recent studies (Miller 1961a, 
1961b, 1962a, 1962b, 1963, Owen 1973, Tavener-Smith 1973, 
Olaloye 1974, Bancroft 1985, 1986b, Bancroft & Wyse Jackson 
1995, Wyse Jackson 1988, Wyse Jackson & Bancroft 1995a) have 
resulted in the description of new taxa, the redescription of previ- 
ously described taxa, and give detailed quantitative and statistical 
analysis of these taxa. While these studies have increased the 
biostratigraphical value of Carboniferous bryozoans from Ireland, 
there is still considerable work to be carried out to assess faunas of 
particular Brigantian stages. 
This present study adds to the taxonomic diversity of bryozoans 
described from Ireland, and shows some similarities at generic level, 
to Lower Carboniferous faunas of the Russian Platform. This paper 
describes 24 bryozoan species of Lower Carboniferous (Viséan, 
Asbian) age from County Fermanagh, Ireland. An unusual nodular 
trepostome and a species of the cystoporate genus Goniocladia, that 
exhibits atypical branching patterns will be described elsewhere. 
PREVIOUS WORK 
The largely silicified fauna from County Fermanagh, dominated by 
bryozoans and brachiopods, has been the subject of several papers: 
Tavener-Smith (1965a) erected the genus Prilofenestella, described 
a species of Minilya (1965b, 1981), noted the occurrence of ovicells 
in Fenestella (1966), described a new species of Polypora (1971), 
and monographed 32 species from eight genera of which three were 
new (1973). Olaloye (1974) examined the acanthocladiid element, 
describing nine species of Penniretepora, five being new. Three new 
fenestrate taxa that were discovered in the Carrick Lough fauna 
during the present study have been described elsewhere (Wyse 
Jackson 1988), and two species of the cryptostome genus / 
Rhabdomeson and the cystoporate taxon Fistulipora incrustans 
(Phillips, 1836) are described more fully by Wyse Jackson & | 
Bancroft (1995a), and Bancroft & Wyse Jackson (1995). 
MATERIAL 
The bryozoans described in this study were collected at two locali- 
ties, Carrick Lough and Sillees River (Fig. 1) from thin beds of pale 
grey and muddy limestones that have been assigned to the upper part | 
of the Glencar Limestone (Brunton & Mason 1979, George er al. | 
1976) (Fig. 2). Nearly 50 kg of rock from Carrick Lough containing | 
both silicified and calcified bryozoan zoaria was processed, and 
additionally many thousands of unsorted etched silicified specimens | 
(from the Brunton and Tavener-Smith collections in the Natural | 
History Museum) and a small number of limestone blocks (from the } 
Mason collection in the Ulster Museum) were examined. A small | 
from drift deposits close to Lough Gara, County Roscommon were | 
also included in this study. | 
Silicified bryozoan colonies were acid-etched from the surround-} 
ing limestones. Calcified bryozoan zoaria were extracted from thei} 
the bryozoans. 
Type and other material from the Griffith Collection in they 
National Museum of Ireland, the Owen Collections in the Manches- 
ter (prefix LL) and Ulster Museums, the Vine Collection in the 
National Museum of Wales, Cardiff (prefix NMW), the Nicholson} 
Collection in Aberdeen University (prefix AUGD), the National} 
Museum of Scotland, Edinburgh (prefix RSM), the Whidborne and} 
