68 
process, D7 neural arch, anterior dorsal rib 
head, rib fragments, r.scapula (proximal) 
D14 neural arch, proximal caudal neural spines 
(3), dorsal rib fragments 
Agastralia, |. fibula proximal fragment 
dorsal rib fragment, |. distalmost rib, 1. pubis 
(distal) 
dorsal rib fragments, gastralia 
Lepidotes teeth & scales 
Iguanodon manual phalanges 
r. angular, dorsal rib heads (4) and shaft 
fragments 
47 
48 
49 
50 
51 
52 
53 
r. surangular, dorsal rib heads (2) and shaft 
fragments 54 
Ce6 neural arch, dorsal rib head fragments 
D6 centrum, D8 centrum, dorsal rib fragments 
Iguanodon dorsal vertebra 
D11 centrum and other centra fragments 
Iguanodon dorsal vertebrae 55 
basal caudal centrum, dorsal rib fragments 
dorsal rib fragments, gastralia, 1. pubis 
fragment, r. femur (distal) 
Lepidotes scales 
Iguanodon supraoccipital, cervical vertebrae, 
A.J. CHARIG AND A.C. MILNER 
indeterminate limb fragments, r. femur (distal) 
D14 centrum, basal caudal centra (2), |. pubis, 
tibia (crushed), astragalus, distal metatarsal 
fragments 
Iguanodon left metatarsals IL, II, IV, phalanges 
(3) 
caudal centrum fragments (2), haemapophyses 
(2) 
* Block not marked on plan 
APPENDIX C: LIST OF CHARACTERS AND DATA-MATRIX, MODIFIED FROM HOLTZ (1994a) 
List of characters employed (the numbers in parentheses at end of each line are Holtz’s original character numbers; the publications that 
follow are those in which these characters were first proposed as synapomorphies). 
SEN A pay ohett 
Pubic plate perforated by pubic fenestra below obturator foramen (1; Rowe 1989, Rowe & Gauthier 1990) 
Distal end of fibula flares to overlap ascending process of astragalus (2; Rowe & Gauthier 1990) 
Sacral ribs fused to centra in adults (3; Rowe & Gauthier 1990) 
Cervical vertebrae with transverse processes strongly backturned and triangular in dorsal view (5; Rowe 1989) 
Pronounced subnarial gap, indicating a possibly mobile premaxillary-maxillary joint (6; Welles 1984, Rowe 1989) 
Metatarsals proximally co-ossified (7; Osmélska 1981, Rowe & Gauthier 1990) 
Crista tibiofibularis with sulcus along medial side of base (8; Rowe 1989, Rowe & Gauthier 1990) 
Axial parapophysis reduced (9; Gauthier 1986) 
Axial diapophysis lost (10; Gauthier 1986) 
Axial pleurocoels lost (11; Gauthier 1986) 
Femoral head directed anteromedially (12; Bonaparte 1991) 
Premaxilla very deep subnarially (13; Bonaparte 1991) 
. Parietal projected dorsally (14; Bonaparte 1991) 
Lower temporal fenestra very large (15; Bonaparte 1991) 
Manual digit III lost (16) 
Manual unguals with pronounced lip on dorsal edge of proximal articulation (17; Currie & Russell 1988) 
Parietals fused with laterosphenoids (18; Barsbold 1983) 
Sternal plates fused (19) 
Pubic boot longer anteriorly than posteriorly (20) 
Pubic boot triangular (apex posterior) in ventral view (21) 
Ilium with preacetabular portion expanded dorsoventrally (22; Gauthier 1986) 
Astragalar condyle with pronounced horizontal groove across anterior face (23; Welles & Long 1974) 
. Supraoccipital crest pronounced (24; Gauthier 1986) 
Premaxillary-maxillary fenestra (25; Osborn 1912) 
Insertion area for pterygoideus muscle below mandibular condyle enlarged (26; Gauthier 1986) 
Lacrimal fenestra present (27; Molnar 1990) 
Orbit key-shaped (28; Gauthier 1986) 
Mandibular fenestra reduced (29; Gauthier 1986) 
Pubic foot projects only posteriorly (30; Gauthier 1986) 
Pelvis opisthopubic (31; Gauthier 1986) 
Pedal digit II longer than IV, closer in length to III (32; Gauthier 1986) 
. Obturator process placed distally (33; Gauthier 1986) 
Ischium not more than two-thirds length of pubis (34; Gauthier 1986) 
. Fourth trochanter lost (35; Gauthier 1986) 
Proximal caudal vertebrae highly modified (neural spine only in 1-9, box-like centra in 1-5, zygapophysial facets vertical) (36, Gauthier 1986) 
Prefrontals extremely reduced or lost (37; Gauthier 1986) 
Frontals separated by an anterior process of parietals (38; Currie 1987) 
Pedal digit Il hyperextensible (39) 
Ulna bowed posteriorly (40; Gauthier 1986) 
Furcula present (41) 
Semilunate carpal present (42; Gauthier 1986) 
Ischial foot lost (43) 
Metacarpal III long and slender (44; Gauthier 1986) 
Ilium with posterodorsal margin curving ventrally in lateral view (45; Gauthier 1986) 
. Coracoid subrectangular (46; Gauthier 1986) 
Chevrons longer than deep (47; Gauthier 1986) 
Sacral vertebrae pleurocoelous (48) 
Dorsal vertebrae procoelous (51) 
Humerus sigmoid in anterior view (52) 
. Quadrate articulation projecting deeply in posteroventral direction (49) 
. Quadrate-quadratojugal articulation mobile (50; Gilmore 1920, Ostrom 1969) 
Ilium with preacetabular portion significantly longer than postacetabular portion (53; Currie & Russell 1988) 
Metatarsals deeper anteroposteriorly than anterolaterally (55; Holtz 1994b) 
Maxillary teeth lost (56) 
Fibula with anterior protuberance below expansion (57; Mader & Bradley 1989) 
. Greater trochanter cleft from head (58; Russell 1969) 
