PALYNOLOGICAL ZONATION OF MID-PALAEOZOIC, NW SPAIN 



(BV) Spore Biozone may be present. The latter zone is based on 

 material from the Anglo-Welsh region (Burgess & Richardson, 

 1991) where the base of the zone, in the type Wenlock area, is 

 recorded in the late hmdgreni Graptolite Biozone (Lower Homerian, 

 Wenlock). 



Scylaspora vetusta - Scylaspora cf. kozlica (Dufka) 

 (V) Spore Assemblage Biozone 



Definition. Based on the co-occurrence of the two nominal taxa 

 with Scylaspora scripta (Burgess & Richardson, 1995). 



Nominal species 



Scylaspora vetusta (Rodriguez) comb. nov. Rodriguez, 1978b- pi 



l,fig8. 

 Scylaspora cf. kozlica (Dufka) comb. nov. Dufka 1995: pi. 3 fios 



lOa-lOb. ' " 



Characteristic species. The species Amhitisporites ahimdo 

 (Rodriguez) comb. nov. and Concentricosisporitcs (?sp.) appear in 

 the upper part of the zone. Within this zone the morphology of 

 Scylaspora vetusta is highly variable. 



Accessory species. Long ranging laevigate species such as 

 Amhitisporites avitus Hoffmeister 1959, A. dihitus (Hoffmeister) 

 Richardson & Lister 1969, A. warringtonii (Richardson & Lister) 

 comb, nov., and Archaeozonotriletes clndus (Cramer) Richardson & 

 Lister 1969 are present, along with murornate Synorisporites sp. 



Remarks. The base of the Scylaspora vetusta Biozone is not seen 

 as both the nomimal species occur in the lowest sample of the La 



119 



Peral section. The upper boundary is defined by the incoming of the 

 nominal species of the succeeding Artemopyra lyrevicosia- 

 Hispanaediscus verrucatus (BV) Biozone. 



Occurence. Strata of the S. vetusta - S. cf. kozlica Spore Biozone 

 are found only in the La Peral section but the nominal species persist 

 into higher zones in other sections. Higher records are discontinuous 

 and may be, at least in part, due to reworking (La Vid Coronaspora 

 reticulata - sanpetrensis and lower Chelinosporu Iwmiesferica (H) 

 Biozones). 



Age. Possibly lower Upper Homerian. Chitinozoa indicate a doubt- 

 ful Llandovery age and may be reworked. 



Artemopyra brevicosta - Hispanaediscm verrucatus 

 (BV) Spore Biozone 



Remarks. Defined in the Anglo- Welsh area (Richardson & 

 McGregor, 1986, Burgess & Richardson. 1995: 26, text-fig. 8) this 

 zone ranges through the middle and upper parts of the Homerian but 

 is unknown in the lower part of the hmdgreni Graptolite Biozone at 

 the base of this stage in the type area. The spore zone in the type area 

 is divided into three subzones, namely: Artemopyra brevicosta. 

 Hispanaediscus lamoutii, and Emphanisporites protophanus (Bur- 

 gess & Richardson, 1 995). In the La Peral sequence the simultaneous 

 appearance of A. brevicosta and H. verrucatus (sample LP4) is 

 followed by the occurrence of Hispanaediscus lamontii in the next 

 higher sample (LP5, 12 m above) and the nominal species persist 

 into the overlying RS Spore Biozone. All of these species are typical 

 of the Homerian, but two (H. kunonti and H. verrucatus) continue 



Spore 

 Zonation 



Loc 



9 



Pri. 



MN 



EC 



Ludf. 



Chitinozoan 

 Zonation 



Loc 



bo 



ele! 



Pri. 



RS 



-?- 



LP 



9 



??ko 



9 



Pri. 



? 



Ludf. 



"?77^?T7 





O O 53 



'^ -a 





P ? is 



~ ~ ,S 



O O t;^ O 





■ S s: 







^ ^ "^ 



5 I ^ a 



■si >; 



5 :« rC 



-S 3 



^ C G 1^ :S ^ ^ 



to U 



6^ 



- 



C^. ■ 0"- 



^Jl* "I* 



Tit ••! 



Ill* 



o 



• • • 



J 



I 





- 



• • 



I 



" Jl 



-IITT 



ill 



Itjtj 



c^- 



■ 



• 



^ Dolomites 

 Limestones 

 ■ Sandstone /Oolites g Sandstone 



I Ferrous Oolites 



I Shale 



• • Phosphatic nodules g Sandstone & Shale 



Al - Aiicyruchiltna Itbyensts 'Late Gorstian / Pndoli 



Ac • Angochirina chtupaciLochkovian 



Eb - Eisenackitina boliernua Lochkovian 



Cb - Cyalhochilina sp, B Late Llandovery 



Me - Margacfutina elegans Mid Pndoli 



Ss- Sphaerochinna sphaerocepiiala Mid Gorstian - early Pridoli 



Pc - PsuedoclathrochiSiiia carmencltui Mid Pndoli 



Uu - Urnochirina urnaPridoli - Earliest Lochkovian 



Arg 15 

 Arg 14 

 Arg 13 

 Aig 12 

 Aig 1TB 

 Arg 11A 

 Arg 10 

 Arg 9 ~ 

 Arg 8 

 Arg 7 

 Arg 5G/6 

 ArgSF 

 Arg 5C 

 Arg 5B 

 Arg 5A 

 Arg 4A 

 Arg 4 

 Arg3 

 Arg 2 

 Arg 1 



MN 

 (na 

 "(aT 

 EC 



RS 



LP 

 9 



Key Cantabrian 

 Chitinozoan taxa 



Ac 



I— ?Eb 



Lochkovian 



Uu/Pc 



/Me Pridoli 



. Ss 



Al Ludfordlan 



Cb 



Fig. 3 Selected spore distribution at Argovejo. Chitinozoan zonation after Verniers el al. ( 1995): bo: bohemica. su: superba. ele: elegans. ko: kosovensis. 



