122 



J.B. RICHARDSON, R.M. RODRIGUEZ AND S.J.E. SUTHERLAND 



with narrow and coarse muri and an irregular reticulum are common 

 accessory spores. Retusotriletes coronadus (Rodriguez, 1983: pi. 8, 

 fig. 1 ) appears in the lower part of the zone. 



Remarks. The upper part of the EC Biozone (Aneurospora Sub- 

 Biozone) is recognisable in England (Hereford and Worcester) in the 

 Raglan Marl Formation below the first appearance of the MN 

 Biozone in the St. Maughans Formation (Richardson, Rodriguez & 

 Sutherland, 2000). In the La Vid section (Fig. 4) the earliest definite 

 specimens of Scylaspora elegans and Aneurospora spp. occur 

 together, consequently the lower part of the EC biozone has not been 

 found there. 



Occurence. The zone is found in all four sections. 



Age. Upper Pffdoli to lowermost Devonian. The appearance of the 

 genus Aneurospora is probably close to the base of the Devonian 

 (Richardson, Ford & Parker 1984; Richardson, Rodriguez & Suther- 

 land, 2000). Chitinozoans in the lower EC Biozone are of Middle 

 Pffdoli age (basal EC, LaPeral,lowerEC,GerasandLa Vid sections). 

 The same chitinozoans indicating a mid-Pfidoli age (Margachitina 

 e/ega«.?jandPfidolitoearliestLochkovianage(t/n7or/2rm!0(/r«a)are 

 both present in the upper part of the EC Biozone (Aneurospora Sub- 

 Biozone) and in the top of the underlying H Biozone. 



Aneurospora spp. (A) Sub-Biozone (Figs 2-5) 

 Reference section for base of sub-biozone. Upper San Pedro 



Formation, La Vid section, Cantabrian Mountains, NW Spain. The 

 lowest occurrence of Aneurospora spp. was found in sample 1 0, 

 c. 106 m above the base of the section. 



Definition. Appearance of the gtrwii, Aneurospora without proxi- 

 mal papillae. Represents the interval between the appearance of 

 Aneurospora and the incoming of Streelispora newportensis of the 

 succeeding zone. 



Accessory species. Spores of the Chelinospora hemiesferica 

 (Cramer & Diez) complex, C. cf. hemiesferica, C. cassicula 

 Richardson & Lister 1969, C. lavidensis sp. nov., C. media sp. nov., 

 Cymbosporites sp. B, and Retusotriletes coronadus s.s. 



Occurence. Found in all four sections in the Cantabrian Mountains. 



Age. Earliest Devonian (Lower Lochkovian), assessment based 

 on comparison with sections in the Lower Old Red Sandstone of 

 England. 



Streelispora newportensis - Emphanisporites 

 micrornatus (MN) Spore Biozone Richardson & 

 McGregor, 1986 (Figs 2-4); Streelispora 

 newportensis - Leonispora argovejae Spore Assem- 

 blage (NA) Sub-Biozone 



Nominal species 



Streelispora newportensis (Chaloner & Streel) Richardson & 

 Lister 1969: pi. 41, figs 3-6 (PI. 7, fig. 8). 





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Me - Margachitina elegans Mid Pridoli 



Ss- Sphaerotiiitin sphaerocepftalaMid Gorstian - Early Pridoli 



S - Spinachittna sp- ''Llandovery 



Pc - Psuedoclalhrochitma carmenchut Mid Pidoll 



Uu • Urnoclutina urna Pridoli - Earliest Lochkovian 



QRed Sandstones 



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□ Sandstone & Shale 



B Sandstone / Oolites 



H Ferrous Oolites 



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LP18 



LP17 



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LP15 



LP14 



LP13 



LP12 



LP11 



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 EC 



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 BV 

 V 



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 S 



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Fig. 5 Selected spore distribution at La Feral. Chitinozoan zonation after Verniers et al. (1995): ele: elegans, ko: kosovensis, ba: barrandei, ph: philipi, el: 

 elongata, ly: lycoperdoides, pa: pachycephala. 



