146 



J.B. RICHARDSON, R.M. RODRIGUEZ AND S.J.E. SUTHERLAND 



Scylaspora cf. kozlica (Dufka) comb. nov. PI. 5, fig. 10 



Cf. 1995 Riigosisporites kozUcus Dufka (pars): 72, pi. 2, figs 15-17. 



Description. Amb circular to subcircular. Equatorial crassitude 

 1-2 |am wide. Trilete mark labrate, c. 4/5 spore radius, lips less than 

 Ipm wide, merge with curvaturae perfectae which form a narrow 

 equatorial crassitude with a thicker inner portion and a narrow (< 0.5 

 |jm), apparently membranous, extension. Proximal surface covered 

 with naiTow irregular rugulae c. 0.5 \xm wide and possible micrograna. 

 Sculpture less distinct near the proximal pole. Distal surface laevigate 

 except for sub-equatorial rugulae with scattered micrograna. 



Dimensions. 3 1-44 nm (30 specimens measured). 



Comparisons. Similar to S. scripta in having radially aligned 

 muri near the equator and distal sculpture, but has more radial 

 geniculate and anastomosing robust muri. S. kozUcus (Dufka) is 

 similar but is distally laevigate and lacks the paired lines of thicken- 

 ing on either side of the Y-mark. S. sp. A (this study) has geniculate 

 muri on the proximal and distal surfaces and is rare. 



Remarks. Scylaspora cf. scripta is similar to forms described 

 from the Homerian and may be reworked. The spores have a mem- 

 branous extension, seen at the equator on some specimens, and may 

 be double-layered with the thin outer part of the exoexine forming 

 the rugulae over the proximal surface. S. vetusta has coarser rugulae 

 but is otherwise closely similar. 



Occurrence. La Vid section, upper RS and lower H Biozones 

 (Upper Ludfordian - Pfidolf). 



Genus CONCENTRICOSISPORITES Rodriguez 1983 



Type species. Concentricosispohtes Sagittarius (Rodriguez) 1983. 



Remarks. Jansonius & Hills (1990: card 4630) considered that 

 the central dark area in the type species may represent an inner body. 



Concentricosisporites agradabilis (Rodriguez) Rodriguez 

 1983 PI. 7, fig. 2 



1 978c Stertozonotriletes agradabilis Rodriguez: 42 1 , pi. 3. figs 7, 



8, 11. 

 1983 Concentricosisporitesagradabilis Rodriguez: 35, pi. 4, figs 



7, 11,13. 



HOLOTYPE AND TYPE LOCALITY. Rodriguez, 1978b: pi. 3, fig. 8: 

 Torrestio, Province of Leon, Cantabrian Mountains, northwest Spain, 

 lower San Pedro Formation. 



Diagnosis. See Rodriguez, 1978c: 421. 



Dimensions. 26-39 pm (18 specimens measured): Rodriguez, 

 1978b, 24-35 Mm. 



Comparisons. This species shows some resemblance to spores of 

 the genus Emphanisporites but the proximal radial ribs are not 

 always distinct and the equatorial crassitude is relatively wide (3-6 

 |am) in relation to the spore radius. 



Occurrence. Lower San Pedro Formation; upper RS and lower 

 H Biozones; sporadically occurs in EC and lower MN Biozones 

 where they may represent reworking. In the higher specimens the 

 distal annulus is lobed and the radial 'muri' more distinct. Upper 

 Ludfordian and Lower Pfidoli. 



Concentricosisporites Sagittarius (Rodriguez) Rodriguez, 

 1983 PI. 7, fig. 1 



1978b Stenozonotriletes Sagittarius Rodriguez: 219, pi. 1, fig. 7. 

 1978c Stenozonotriletes Sagittarius Rodriguez: 421, pi. 2, figs 13, 



14. 

 1 983 Concentricosisporites Sagittarius (Rodriguez); Rodriguez: 



36,pl. 3,fig. 21. 

 1995 Concentricosisporites Sagittarius Rodriguez; Burgess & 



Richardson: 17, pi. 6, figs 14. 15. 



Holotype. Rodriguez, 1978b: pi. 1, fig. 7. 



Diagnosis. See Rodriguez, 1978b: 219. 



Dimensions. 29^4 pm (10 specimens measured); 25-40 |am 

 (Rodriguez, 1978). 



Occurrence. San Pedro Formation, Upper RS & H Biozones La 

 Vid, upper RS & lower H Biozones La Peral, lower MN Biozone 

 Geras (? reworked). Upper Ludfordian and Lower Pfidoli. 



Genus IBEROESPORA Cramer & Diez 1975 



Type species. Iberoespora cantabrica Cramer & Diez 1975. 



Iberoespora cantabrica Cramer & Diez 1975 



PI. 6, fig. 4; PI. 7, fig. 4 



Holotype. Cramer & Diez 1975: pi. 2, fig. 24. 



Diagnosis. See Cramer & Diez 1975: 339, pi. 2, figs, 24, 26-28, 

 30.31. 



Dimensions. 30 to 45 pm (Cramer & Diez, 1975);27to42|jm(40 

 specimens measured). 



PLATE 7 



Fig. 1 Concentricosisporites Sagittarius (Rodriguez) Rodriguez 1983. FM 1514, Ger92/15a (475) 1, 044 1032. 



Fig. 2 Concentricosisporites agradabilis (Rodriguez) Rodriguez 1983. FM 1515. LV92/13 (495) 2. 098 0982; 2a, proximal focus; 2b, distal focus. 



showing annulus. 

 Fig.3 Iberoespora glabella! CrameT &D\ez 1975. FM 1516, Argovejo 92/12 (510) 3, 1390874. 

 Fig. 4 Iberoespora cantabrica Cramer & Diez 1975. FM 1517, Arg92/12 (510) 3, 212 0962; 4a, 4b, showing verrucate lips, distal geniculate muri, and 



inter-radial thickenings; 4a, x 1500; 4c, median focus showing margin of the central body invaginated. 

 Fig.5 Uonispora argovejae Cramer & Diez 1975. FM 1519. LV92/13 (D3) 1.051 0913. 



Iberoespora mariae Rodriguez 1983. FM 1518, LP92/20 (D8) 2, Geras 92/1 1 (D8) 1, 030 0969 (see Fig.2, sample GER 1 1 ). 

 Aneurospora sp. Tripapillate form. FM 1520, Ger92/14 (468) 7, 132 0975; 7a, proximal focus; 7b, proximal focus, showing curvaturae perfectae. 

 Street ispora newportensis (Chaloner & Streel) Richardson & Lister 1969. FM 1521, LV13 (D3) 1, 082 0986; 8a, 8b. x 1500. 

 Aneurospora richardsonii (Rodriguez) comb, nov., FM 1522, Gerl4 92/14 (468) 7, 135 0838; 9b, shows sculpture in detail, x 1500. 

 Chelinospora sanpetrensis (Rodriguez) comb. nov. FM 1523, Arg92/5a (473) 3, 123 1092; 10a, distal focus showing foveolate exine; 10b, with 



Fig. 6 

 Fig. 7 

 Fig. 8 

 Fig. 9 

 Fig. 10 



narrow arcuate fold around proximal hilum, x 1 500. 

 All figs X 1000, unless stated otherwise. 



