PALYNOLOGICAL ZONATION OF MID-PALAEOZOIC, NW SPAIN 



Comparison. Distally foveolate specimens with a laevigate 

 crassitude are referred to as Iberoespora glabella? {P\. 6, fig. 5: PL 7. 

 fig. 3). The original illustrations (Cramer & Diez. 1975: pi. 2, figs 22, 

 29) do not show the distal surface clearly. 



Remarks. Cramer & Diez do not explain how they differentiate /. 

 guzmani from /. cantabrica. Both species have the same structure, 

 radial muri on the crassitude, crenulate lips, a distal sculpture of 

 reticulate to convolute muri, and thickenings in the inter-radial areas. 

 However, the size range given for /. guzmani is only 10-20 |am 

 whereas that for /. cantabrica is 30-45 |jm. The specimens in this 

 study are all larger than 26 |jm. 



Occurrence. Upper San Pedro Formation in Argovejo, Geras 

 and La Vid sections in the Province of Leon and La Peral section in 

 the Province of Asturias, upper EC - lower MN Biozones. 



Genus LEONISPORA Cramer & Diez 1975 

 Type species. Leonispora argovejae Cramer & Diez 1975. 



Leonispora argovejae Cramer & Diez 1975 PI. 7, fig. 5 



Holotype. Cramer & Diez 1975, pi. 1, fig. 3. 



Description. Cramer & Diez (1975: 342). 



Dimensions. 25-35 |am (ibid. 1975); 30-38 \xm (14 specimens 

 measured, present study). 



Comparisons. The genus is closely similar in structure to 

 Streelispora but is laevigate. 



Occurrence. Upper San Pedro Formation, Geras, La Vid sec- 

 tions; MN Miospore Biozone, NA Subzone. 



Genus STREELISPORA (Richardson & Lister) Richardson, 

 Streel, Hassan & Steemans 1982 



Type species. Streelispora newportensis (Chaloner & Streel); 

 Richardson & Lister 1969. 



149 



Dimensions. 28-39 pm (13 specimens measured). 17^8 pm (80 

 specimens measured) Richardson & Lister 1969. 



Occurrence. Upper San Pedro Formation, Argovejo, Geras. La 

 Vid, MN Biozone, NA Sub-Biozone. 



Genus ANEUROSPORA Streel 1964 

 Type species. Aneurospora goensis Stred 1964 



Aneurospora richardsonii (Rodriguez) comb. nov. 



Pi. 4, fig. 1 ; PI. 7, fig. 9 



Basionym. Streelispora richardsonii Rodriguez, 1983: 50, pi. 8. 

 figs 6, 7, pi. 9, fig. 3. 



1 978c Geminospora spinosa Allen; Rodriguez: 4 1 6, pi. 5, figs 7, 8. 

 1983 Streelispora richardsonii Rodriguez: 50, pi. 8, figs 6, 7, pi. 

 9, fig. 3. 



Holotype and locus typicus. See Rodriguez, 1983: 50. pi. 8. 

 fig. 3. 



Emended diagnosis. An Aneurospora with no proximal papillae, 

 distal sculpture consists of spaced slender spines (fimbriae), ele- 

 ments dominantly parallel-sided with blunt, rounded apices. 



Comparison. The slender fimbriae distinguish this species. 

 Anapiculatisporites raistrickiaeformis Schultz (1968) has coarser, 

 more irregular elements (see also Steemans 1989). 



Dimensions. 38 to 50 jjm (Rodriguez, 1983); 32 to 47 pm (28 

 specimens measured). 



Occurrence. Lower part of lower MN Biozone, NA Subzone, 

 Argovejo, Geras and La Vid sections. 



Remarks. The species Aneurospora richardsonii lacks the proxi- 

 mal papillae and associated tangential folds and is herein transferred 

 to the genus Aneurospora. Tangential folds around the proximal 

 papillae distinguish Streelispora and Leonispora from tripapillate 

 species oi Aneurospora (eg. PI. 7, fig. 7). 



Streelispora newportensis (Chaloner & Streel); Richardson 

 & Lister 1969 PL 7, fig. 8 



1968 Granulatisporites newportensis Chaloner & Streel: 92, pi. 

 19, figs 7, 8, 11. 



1 969 Streelispora newportensis (Chaloner & Streel); Richardson 

 & Lister: 230-31, pi. 41, figs 3-6. 



Holotype. Chaloner & Streel 1968: 92, pi. 19, figs 7, 8, 1 1. 



Genus CORONASPORA Rodriguez 1979 emend. 



1978b Coronaspora niariae Rodriguez: 218. pi. 1, figs 1-3. 

 1979 Coronaspora mariae Rodriguez: 232. 



Type SPECIES. Coronaspora mariae Rodvigutz, 1978. 



Holotype. Coronaspora mariae Rodriguez. 1978b: pi. 1. figs 1, 

 2. 



PLATE 8 



Figs 1-4 Coronaspora cromatica (Rodriguez) emend. 1, FM 1524; la, showing proximal kyrtome and broad verrucate lips. lb. intermediate focus. Ic, 



distal view showing annulus and sculpture of sinuous muri and rounded verrucae; Ger92/5 (542) 2, 098 1006; 2, FM 1525; 2a-c, proximal to distal levels 



of focus; 2a with narrow lips and distinct kyrtome; 2b relatively narrow crassitude; 2c, distal sculpture of small interconnected narrow verrucae Arg92/5a. 



(473) 3, 023 0983; 3, FM 1526, proximal view, Arg92/5a (473) 3, 139 1 133; 4, FM 1527, proximal view, specimen with inter-radial thickenings on the 



kyrtome, Arg92/5a (473) 3. 099 0938. 

 Figs 5, 6 Coronaspora reticulata sp. nov, 5, FM 1528, proximal focus showing kyrtome with wide inter-radial thickenings. LV92/7b (D3) 3. 156 0917; 6a, 



b, FM 1491 (holotype). showing distal reticulum (the radial extremities are visible in Fig. 6a). LV92/7b (03) 3. 043 0965. 

 Fig. 7 Coronaspora siihornata (Cramer & Diez) comb. nov. FM 1 529. specimen showing narrow Y-folds. equatorial crassitude, distal annulus and 



irregular distal thickening; Ger92/9 (456) 1, 035 0472. 

 Fig. 8 Chelinospora canistrata sp. nov. FM 1492 (holotype), Ger92/5 (542) 2. 083 1 024. 

 Fig. 9 Coronaspora infraornata (Rodriguez) comb, nov, FM 1530. showing curvatural crassitude, distal annulus and distal polar thickening at two focal 



levels, Ger92/9 (456) 1. 066 0942. 



All tigs X 1000. 



